scholarly journals Historical biogeography of the leopard (Panthera pardus) and its extinct Eurasian populations

2018 ◽  
Author(s):  
Johanna L.A. Paijmans ◽  
Axel Barlow ◽  
Daniel W. Förster ◽  
Kirstin Henneberger ◽  
Matthias Meyer ◽  
...  
Keyword(s):  
Late Pleistocene ◽  
Dna Sequences ◽  
Sister Group ◽  
Historical Biogeography ◽  
Early Pleistocene ◽  
Middle Pleistocene ◽  
Recent Common Ancestor ◽  
Panthera Pardus ◽  
Phylogenetic Placement ◽  
Out Of Africa

AbstractBackgroundResolving the historical biogeography of the leopard(Panthera pardus)is a complex issue, because patterns inferred from fossils and from molecular data lack congruence. Fossil evidence supports an African origin, and suggests that leopards were already present in Eurasia during the Early Pleistocene. Analysis of DNA sequences however, suggests a more recent, Middle Pleistocene shared ancestry of Asian and African leopards. These contrasting patterns led researchers to propose a two-stage hypothesis of leopard dispersal out of Africa: an initial Early Pleistocene colonisation of Asia and a subsequent replacement by a second colonisation wave during the Middle Pleistocene. The status of Late Pleistocene European leopards within this scenario is unclear: were these populations remnants of the first dispersal, or do the last surviving European leopards share more recent ancestry with their African counterparts?ResultsIn this study, we generate and analyse mitogenome sequences from historical samples that span the entire modern leopard distribution, as well as from Late Pleistocene remains. We find a deep bifurcation between African and Eurasian mitochondrial lineages (∼710 Ka), with the European ancient samples as sister to all Asian lineages (∼483 Ka). The modern and historical mainland Asian lineages share a relatively recent common ancestor (∼122 Ka), and we find one Javan sample nested within these.ConclusionsThe phylogenetic placement of the ancient European leopard as sister group to Asian leopards suggests that these populations originate from the same out-of-Africa dispersal which founded the Asian lineages. The coalescence time found for the mitochondrial lineages aligns well with the earliest undisputed fossils in Eurasia, and thus encourages a re-evaluation of the identification of the much older putative leopard fossils from the region. The relatively recent ancestry of all mainland Asian leopard lineages suggests that these populations underwent a severe population bottleneck during the Pleistocene. Finally, although only based on a single sample, the unexpected phylogenetic placement of the Javan leopard could be interpreted as evidence for exchange of mitochondrial lineages between Java and mainland Asia, calling for further investigation into the evolutionary history of this subspecies.

Bayesian analysis of sea-level sensitivity to CO2 forcing across the mid-Pleistocene transition: possible implications for early-Pleistocene ice-sheet extent

2021 ◽  
Author(s):  
Parker Liautaud ◽  
Peter Huybers
Keyword(s):  
Energy Balance ◽  
Bayesian Analysis ◽  
Sea Level ◽  
Late Pleistocene ◽  
Radiative Forcing ◽  
Ice Sheets ◽  
Ice Sheet ◽  
Early Pleistocene ◽  
Middle Pleistocene ◽  
Model Parameters

<p><span>Foregoing studies have found that sea-level transitioned to becoming approximately twice as sensitive to CO</span><span><sub>2</sub></span><span> radiative forcing between the early and late Pleistocene (Chalk et al., 2017; Dyez et al., 2018). In this study we analyze the relationships among sea-level, orbital variations, and CO</span><span><sub>2</sub></span><span> observations in a time-dependent, zonally-averaged energy balance model having a simple ice sheet. Probability distributions for model parameters are inferred using a hierarchical Bayesian method representing model and data uncertainties, including those arising from uncertain geological age models. We find that well-established nonlinearities in the climate system can explain sea-level becoming 2.5x (2.1x - 4.5x) more sensitive to radiative forcing between 2 and 0 Ma. Denial-of-mechanism experiments show that the increase in sensitivity is diminished by 36% (31% - 39%) if omitting geometric effects associated with thickening of a larger ice sheet, by 81% (73% - 92%) if omitting the ice-albedo feedback, and by more than 96% (93% - 98%) if omitting both. We also show that prescribing a fixed sea-level age model leads to different inferences of ice-sheet dimension, planetary albedo, and lags in the response to radiative forcing than if using a more complete approach in which sea-level ages are jointly inferred with model physics. Consistency of the model ice-sheet with geologic constraints on the southern terminus of the Laurentide ice sheet can be obtained by prescribing lower basal shear stress during the early Pleistocene, but such more-expansive ice sheets imply lower CO</span><span><sub>2</sub></span><span> levels than would an ice-sheet having the same aspect ratio as in the late Pleistocene, exacerbating disagreements with </span><span>𝛿</span><span><sup>11</sup></span><span>B-derived CO</span><span><sub>2</sub></span><span> estimates. These results raise a number of possibilities, including that (1) geologic evidence for expansive early-Pleistocene ice sheets represents only intermittent and spatially-limited ice-margin advances, (2) </span><span>𝛿</span><span><sup>11</sup></span><span>B-derived CO</span><span><sub>2</sub></span><span> reconstructions are biased high, or (3) that another component of the global energy balance system, such as the average ice albedo or a process not included in our model, also changed through the middle Pleistocene. Future work will seek to better constrain early-Pleistocene CO</span><span><sub>2</sub></span><span> levels by way of a more complete incorporation of proxy uncertainties and biases into the Bayesian analysis.</span></p>

scholarly journals Phylogenetic relationships, origin and historical biogeography of the genus Sprattus (Clupeiformes: Clupeidae)

PeerJ ◽  
2021 ◽  
Vol 9 ◽  
pp. e11737
Author(s):  
Cristian B. Canales-Aguirre ◽  
Peter A. Ritchie ◽  
Sebastián Hernández ◽  
Victoria Herrera-Yañez ◽  
Sandra Ferrada Fuentes ◽  
...  
Keyword(s):  
Southern Hemisphere ◽  
Northern Hemisphere ◽  
Dna Sequences ◽  
Phylogenetic Analyses ◽  
Historical Biogeography ◽  
Ancestral Population ◽  
Recent Common Ancestor ◽  
Most Recent Common Ancestor ◽  
The Tropics ◽  
Two Populations

The genus Sprattus comprises five species of marine pelagic fishes distributed worldwide in antitropical, temperate waters. Their distribution suggests an ancient origin during a cold period of the earth’s history. In this study, we evaluated this hypothesis and corroborated the non-monophyly of the genus Sprattus, using a phylogenetic approach based on DNA sequences of five mitochondrial genome regions. Sprattus sprattus is more closely related to members of the genus Clupea than to other Sprattus species. We also investigated the historical biogeography of the genus, with the phylogenetic tree showing two well-supported clades corresponding to the species distribution in each hemisphere. Time-calibrated phylogenetic analyses showed that an ancient divergence between Northern and Southern Hemispheres occurred at 55.8 MYBP, followed by a diversification in the Oligocene epoch in the Northern Hemisphere clade (33.8 MYBP) and a more recent diversification in the Southern Hemisphere clade (34.2 MYBP). Historical biogeography analyses indicated that the most recent common ancestor (MRCA) likely inhabited the Atlantic Ocean in the Southern Hemisphere. These results suggest that the ancestral population of the MRCA diverged in two populations, one was dispersed to the Northern Hemisphere and the other across the Southern Hemisphere. Given that the Eocene was the warmest epoch since the Paleogene, the ancestral populations would have crossed the tropics through deeper cooler waters, as proposed by the isothermal submergence hypothesis. The non-monophyly confirmed for the genus Sprattus indicates that its systematics should be re-evaluated.

Biostratigraphic Correlation of Pleistocene Marine Deposits and Sea Levels, Atlantic Coastal Plain of the Southeastern United States

1980 ◽  
Vol 13 (2) ◽  
pp. 213-229 ◽  
Author(s):  
Thomas M. Cronin
Keyword(s):  
North Carolina ◽  
South Carolina ◽  
Sea Level ◽  
Late Pleistocene ◽  
Coastal Plain ◽  
Atlantic Coastal Plain ◽  
Early Pleistocene ◽  
Middle Pleistocene ◽  
Sea Levels ◽  
Atlantic Coastal

AbstractMarine ostracodes from 50 localities were studied to determine the age and elevation of Pleistocene sea levels in the Atlantic coastal plain from Maryland to northern Florida. Using ostracode taxon and concurrent ranges, published planktic biostratigraphic, paleomagnetic, and radiometric data, ostracode assemblage zones representing early (1.8-1.0 my), middle (0.7-0.4 my), and late (0.3-0.01 my) Pleistocene deposition were recognized and used as a basis for correlation. Ostracode biofacies signifying lagoonal, oyster bank, estuarine, open sound, and inner sublittoral environments provided estimated ranges of paleodepths for each locality. From these data the following minimum and maximum Pleistocene sea-level estimates were determined for the southeastern coastal plain: late Pleistocene, 2–10 m from Maryland to northern Florida; middle Pleistocene, 6–15 m in northern South Carolina; early Pleistocene, 4–22 m in central North Carolina, 13–35 m in southern North Carolina, and 6–27 m in South Carolina. Climatically induced glacio-eustatic sea-level fluctuations adequately account for the late Pleistocene sea-level data, but other factors, possibly differential crustal uplift, may have complicated the early Pleistocene record.

scholarly journals Long-term landscape evolution of the Molise sector of the central-southern Apennines, Italy

2017 ◽  
Vol 68 (1) ◽  
pp. 29-42 ◽  
Author(s):  
Vincenzo Amato ◽  
Pietro P.C. Aucelli ◽  
Vito Bracone ◽  
Massimo Cesarano ◽  
Carmen Maria Rosskopf
Keyword(s):  
Late Pleistocene ◽  
Landscape Evolution ◽  
Temporal Distribution ◽  
Structural Data ◽  
Early Pleistocene ◽  
Middle Pleistocene ◽  
Southern Apennines ◽  
Geomorphological Evolution ◽  
Present Setting

AbstractThis paper concerns the reconstruction of the main stages of the long-term landscape evolution of the Molise portion of the central-southern Apennines along a transect divided into three sectors (SW, Central and NE). Analysis mainly focused on geomorphological, stratigraphical and structural data supported by chronological constraints, coming from an overall review of past literature and several studies carried out by the authors of the paper during the last 20 years. The results obtained allowed the elaboration of a conceptual model of the long-term evolution of the Molise sector of the central-southern Apennines. Starting from the Pliocene, the emersion of the Molise area occurred gradually from SW to NE, allowing a polycyclic landscape to evolve under the major controls first of compression then transtensional to extensional tectonics as well as climatic variations. Principal markers of the Quaternary geomorphological evolution of the Molise area are represented by the infill successions of the intermontane tectonic depressions located in its internal, SW sector and by four orders of palaeosurfaces that developed between the Early Pleistocene and the beginning of the Late Pleistocene across the region. These markers testify to the alternation of phases of substantial tectonic stability and uplift whose spatial-temporal distribution could be assessed along the investigated transect. Results highlight that the most important stages of landscape evolution occurred during the Early and Middle Pleistocene. At the beginning of the Late Pleistocene, the Molise sector of the Apennine chain had already reached its present setting and further landscape evolution occurred under the major control of climate and land-use.

scholarly journals ABOUT THE REASONS OF CLIMATE WARMING BASED ON STUDYING THE HISTORY OF QUATERNARY GLACIOSES OF THE CAUCASUS (ON THE EXAMPLE OF THE INTERDURCHIE TEREK AND THE ANDIAN KOISU)

2020 ◽  
Vol 12 (3) ◽  
pp. 461-471
Author(s):  
Valeriy DOTSENKO ◽  
◽  
Ibragim KERIMOV ◽  
Keyword(s):  
Climate Change ◽  
Late Pleistocene ◽  
Volcanic Activity ◽  
Late Holocene ◽  
Early Pleistocene ◽  
Middle Pleistocene ◽  
The Caucasus ◽  
Late Pliocene ◽  
Climate Fluctuations ◽  
Greater Caucasus

The Greater Caucasus experienced repeated glaciation during the Quaternary (early, middle, upper Pleistocene, late Glacial, and late Holocene), which occurred under changing climatic conditions and differentiated tectonic movements. These glaciations, of course, are associated with changes in terrain, the formation of new deposits, transgressions and regressions of the Caspian Sea, changes in vegetation and soil types, so the problem of glaciation affects all earth Sciences to varying degrees. The study of Quaternary glaciation, especially Holocene glaciation, is currently relevant for understanding climate change. Against the background of significant climate fluctuations within the epochs of glaciation, there are smaller cooling phases that cause the temporary onset of glaciers. Short-term climate fluctuations are manifested in oscillations – minor fluctuations in the languages of glaciers. All this indicates that the climate undergoes significant changes in a short time, which are reflected in the morphosculpture of the terrain, the latest deposits and modern precipitation. Glaciation of the Greater Caucasus in the Prikazbeksky region reached its maximum in the middle Pleistocene,when glaciers went far into the Ossetian basin. All these traces have been preserved due to the lower capacity of the Chanty-Argun glacier and its fluvioglacial flow, which developed during the late Pleistocene epoch. Volcanic activity, especially active in the late Pliocene and continuing up to the present time, is associated with the late horn stage of development of the Caucasus. The formation of the Rukhs-Dzuar molass formation more than 2 km thick in the late Pleistocene in the Ossetian basin of the Tersky-Caspian flexure is associated with the activity of volcanoes in the Kazbek volcanic region. In the early Pleistocene, volcanic activity on the BC decreased significantly. The most intense outbreak of volcanism in the Kazbek and Elbrus volcanic regions occurred at the beginning of the late Pleistocene, which roughly coincided with the maximum phase of the late Pleistocene (Bezengian) glaciation. Then, in the second half of the late Pleistocene, volcanic activity was manifested on the mount Kazbek. The last outbreak of volcanic activity occurred in the Holocene no more than 2-3 thousand years ago. Fresh lavas are available on Elbrus, Kazbek, in the Terek valley near villages. Sioni and on the Kel volcanic plateau. Fumarolic activity still continues on Elbrus. Thus, in the Kazbek region, eruptions occurred from the late Pliocene to the late Holocene inclusive. Keywords: Pleistocene, Holocene, glaciation stages, nival-glacial processes, causes of glaciations, climate change, anthropogenic factors, natural factors, Earth degassing, magmatogenic degassing branch, seismotectonic degassing branch, greenhouse gases, newest geodynamics, volcanism, mud regimes, volcanism, methane hydrates, land degradation, water reclamation.

Increased sea-level sensitivity to CO2 forcing across the Middle Pleistocene Transition from ice-albedo and ice-volume nonlinearities

2021 ◽  
pp. 1
Author(s):  
Parker Liautaud ◽  
Peter Huybers
Keyword(s):  
Sea Level ◽  
Late Pleistocene ◽  
Radiative Forcing ◽  
Ice Sheets ◽  
Ice Sheet ◽  
Early Pleistocene ◽  
Balance Model ◽  
Middle Pleistocene ◽  
Model Parameters ◽  
The Past

AbstractProxy reconstructions indicate that sea level responded more sensitively to CO2 radiative forcing in the late Pleistocene than in the early Pleistocene, a transition that was proposed to arise from changes in ice-sheet dynamics. In this study we analyse the links between sea level, orbital variations, and CO2 using an energy-balance model having a simple ice sheet. Model parameters, including for age models, are inferred over the late Pleistocene using a hierarchical Bayesian method, and the inferred relationships are used to evaluate CO2 levels over the past 2 My in relation to sea level. Early-Pleistocene model CO2 averages 246 ppm (244 ppm - 249 ppm 95% c.i.) across 2-1 Ma and indicates that sea level was less sensitive to radiative forcing than in the late Pleistocene, consistent with foregoing δ11B-derived estimates. Weaker early-Pleistocene sea-level sensitivity originates from a weaker ice-albedo feedback and the fact that smaller ice sheets are thinner, absent changes over time in model equations or parameters. An alternative scenario involving thin and expansive early-Pleistocene ice sheets, in accord with some lines of geologic evidence, implies 15 ppm lower average CO2 or ~10-15 m higher average sea level during the early Pleistocene relative to the original scenario. Our results do not rule out dynamical transitions during the middle Pleistocene, but indicate that variations in the sea-level response to CO2 forcing over the past 2 My can be explained on the basis of nonlinearities associated with ice-albedo feedbacks and ice-sheet geometry that are consistently present across this interval.

Early Pleistocene−to−present paleoclimate archive for the American Southwest from Stoneman Lake, Arizona, USA

2021 ◽  
Author(s):  
Spencer E. Staley ◽  
Peter J. Fawcett ◽  
R. Scott Anderson ◽  
Gonzalo Jiménez-Moreno
Keyword(s):  
Late Pleistocene ◽  
Climate Model ◽  
Volcanic Eruptions ◽  
American Southwest ◽  
Early Pleistocene ◽  
Middle Pleistocene ◽  
Future Climate Change ◽  
Radiocarbon Dates ◽  
Middle And Late Pleistocene ◽  
Natural Climate

Long, continuous records of terrestrial paleoclimate offer insights into natural climate variability and provide context for geomorphological studies, climate model reconstructions, and predictions of future climate change. STL14 is an 80 m lacustrine sediment core that archives paleoenvironmental changes at Stoneman Lake, Coconino County, Arizona, from the early Pleistocene (ca. 1.3 Ma) to present. Full-core sedimentology was analyzed using smear slides and core face observations. Lithofacies strongly correlate with wet bulk density and bulk magnetic susceptibility (MS), and these data resemble a sawtooth pattern characteristic of glacial-interglacial climate cycles. A linkage between deep to shallow lake depth transitions and glacial terminations is supported by an age model that incorporates accelerator mass spectrometry radiocarbon dates and tephrochronology of ashes from the Lava Creek B and multiple Long Valley, California, volcanic eruptions. We correlated middle and late Pleistocene glacial maxima to deep lake deposits defined by well-preserved bedding, increased biosilica, boreal pollen taxa (i.e., Picea), and lower density and MS. Interglacial periods are associated with shallow-water deposits characterized by banded-to-massive siliciclastic material, some authigenic calcite, the alga Phacotus, and higher density and MS. Prior to the marine isotope stage (MIS) 24−22 interval, smaller-amplitude changes in the lake environment suggest milder glacial conditions compared to those of the middle and late Pleistocene. Thus, abrupt intensification of glacial conditions may have occurred ca. 900 ka in the American Southwest, mirroring a global characteristic of the mid-Pleistocene transition. The STL14 record suggests that lake environments throughout the history of this small (3.5 km2), internally drained, basaltic catchment are sensitive to the regional hydrologic balance, which, at orbital time scales, is largely influenced by the northern cryosphere and associated changes in atmospheric circulation. The predominance of quartz in sediment throughout the record indicates significant eolian inputs. Few paleoclimate records from this region extend beyond the last glacial cycle, let alone the middle Pleistocene, making STL14 a valuable resource for studying environmental responses to a range of natural climate states and transitions throughout much of the Quaternary.

Stratigraphy and structure of Pleistocene collapse in the Regina Low, Saskatchewan, Canada

2002 ◽  
Vol 39 (9) ◽  
pp. 1411-1423 ◽  
Author(s):  
E A Christiansen ◽  
E Karl Sauer
Keyword(s):  
Late Pleistocene ◽  
Middle Devonian ◽  
Upper Cretaceous ◽  
Early Pleistocene ◽  
Middle Pleistocene ◽  
Pierre Shale ◽  
Hydraulic Gradients ◽  
Group A ◽  
Structural Closure

The Regina Low is a collapse structure, formed as a result of dissolution of salt from the Middle Devonian Prairie Evaporite Formation. In this study, collapse has affected the Upper Cretaceous Pierre Shale and the Ardkenneth and Snakebite members of the Bearpaw Formation of the Montana Group; the Mennon, Dundurn, and Warman formations of the Sutherland Group; and the Floral and Battleford formations of the Saskatoon Group. A structural closure of 125–175 m approximates the thickness of the Middle Devonian Prairie Evaporite Formation. In the Early Pleistocene, about 54 m of collapse took place in eastern Regina accounting for the preservation of Snakebite Member. Major collapses of about 58 and 86 m took place in northern Regina during deposition of Middle Pleistocene, pre-Illinoian Dundurn Formation. The final major collapse of about 127 m took place in northeastern Regina between deposition of the lower and upper tills of the Floral Formation. Eighty-seven metres of this collapse took place during deposition of the Late Pleistocene, interglacial, Sangamon Pasqua Member of the Floral Formation. The Pleistocene fill in the Regina Low collapse structure suggests that collapse took place when dissolution of salt from the Prairie Evaporite Formation was accelerated by high hydraulic gradients created by the surcharge pressures of the glaciers.

scholarly journals Early Pleistocene enamel proteome sequences from Dmanisi resolve Stephanorhinus phylogeny

2018 ◽  
Author(s):  
Enrico Cappellini ◽  
Frido Welker ◽  
Luca Pandolfi ◽  
Jazmin Ramos Madrigal ◽  
Anna K. Fotakis ◽  
...  
Keyword(s):  
Phylogenetic Analyses ◽  
Dental Enamel ◽  
Sister Group ◽  
Direct Access ◽  
Early Pleistocene ◽  
Middle Pleistocene ◽  
Time Range ◽  
Molecular Evidence ◽  
Type I ◽  
Taxonomic Assignment

ABSTRACTAncient DNA (aDNA) sequencing has enabled unprecedented reconstruction of speciation, migration, and admixture events for extinct taxa1. Outside the permafrost, however, irreversible aDNA post-mortem degradation2 has so far limited aDNA recovery within the ˜0.5 million years (Ma) time range3. Tandem mass spectrometry (MS)-based collagen type I (COL1) sequencing provides direct access to older biomolecular information4, though with limited phylogenetic use. In the absence of molecular evidence, the speciation of several Early and Middle Pleistocene extinct species remain contentious. In this study, we address the phylogenetic relationships of the Eurasian Pleistocene Rhinocerotidae5-7 using ˜1.77 million years (Ma) old dental enamel proteome sequences of a Stephanorhinus specimen from the Dmanisi archaeological site in Georgia (South Caucasus)8. Molecular phylogenetic analyses place the Dmanisi Stephanorhinus as a sister group to the woolly (Coelodonta antiquitatis) and Merck’s rhinoceros (S. kirchbergensis) clade. We show that Coelodonta evolved from an early Stephanorhinus lineage and that this genus includes at least two distinct evolutionary lines. As such, the genus Stephanorhinus is currently paraphyletic and its systematic revision is therefore needed. We demonstrate that Early Pleistocene dental enamel proteome sequencing overcomes the limits of ancient collagen- and aDNA-based phylogenetic inference, and also provides additional information about the sex and taxonomic assignment of the specimens analysed. Dental enamel, the hardest tissue in vertebrates, is highly abundant in the fossil record. Our findings reveal that palaeoproteomic investigation of this material can push biomolecular investigation further back into the Early Pleistocene.

scholarly journals Pliocene–Pleistocene sedimentary and geomorphologic development of the Vasilikos river catchment, S Cyprus, in relation to uplift of the Troodos ophiolite and climate-related changes

2019 ◽  
Vol 157 (4) ◽  
pp. 573-602 ◽  
Author(s):  
Hannah Murray ◽  
Alastair HF Robertson
Keyword(s):  
Late Pleistocene ◽  
Close Correlation ◽  
Sediment Supply ◽  
Early Pleistocene ◽  
Middle Pleistocene ◽  
Terra Rossa ◽  
River Catchment ◽  
Fluvial Terraces ◽  
Surface Uplift ◽  
Fluvial Sedimentation

AbstractThe Pleistocene development of the Vasilikos River exemplifies the interaction of focused, tectonically induced surface uplift and climate-influenced changes. The resulting sediments are well exposed in Vasilikos Quarry and in the main river catchment further east. An important erosional surface incises the highest-level (oldest) fluvial conglomerates, down into Late Pliocene – Early Pleistocene open-marine mudrocks (Nicosia Formation), allowing integration with the circum-Cyprus sedimentary–geomorphic development (F1–F4 stages). To determine where the quarry deposits lie in relation to the Vasilikos river catchment, the fluvial deposits were mapped and valley profiles were constructed, revealing four main episodes, each associated with incision and distinctive fluvial deposition. Source lithology strongly influenced channel morphology, infill and adjacent slope-sediment (colluvium) composition. Palaeosols, particularly red-brown terra rossa, developed on abandoned fluvial terraces and adjacent hillslopes, especially overlying F3 surfaces. The combined evidence allows close correlation of the Vasilikos river and quarry deposits. Relatively coarse (chalky conglomerate/breccia) and fine-grained colluvium (calcareous silt – Cyprus harvara) developed especially on lower hillslopes following incision (mainly above F2 and F3 surfaces). Based on regional comparisons, overall sediment aggregation ended during the Early Pleistocene. The F1–F2 surfaces and deposits are inferred to be Middle Pleistocene, the F3 ones later Middle Pleistocene and the F4 ones near the Middle–Late Pleistocene boundary. Geomorphology and deposition were tectonically forced during strong, focused Early–Middle Pleistocene surface uplift. Coarse clastic ruff-off and palaeosol development (terra rossa) and related sediment aggradation are inferred to have increased during warm, humid periods. Late Pleistocene geomorphology and deposition were more influenced by climatic change, with semi-perennial streamflow, rapid sediment aggradation and palaeosol (terra rossa) development during warm, humid periods (interglacials). Cooler (glacial) periods enhanced fluvial-incision, sediment-bypassing and hillslope colluvial processes (e.g. frost shattering, downslope creep and mass flow) when sediment transport (bypassing) exceeded sediment supply. Neotectonic faulting affected the catchment but did not greatly affect geomorphology or sediment supply. Although climate / climate change (and eustatic sea-level change) had an important influence, tectonics is interpreted as the fundamental driver of geomorphological development and fluvial sedimentation, with implications for other areas, regionally to globally.

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