Morphological phylogeny of the Australian genera of the bee fly subfamily Bombyliinae (Diptera : Bombyliidae) with description of four new genera

2018 ◽  
Vol 32 (2) ◽  
pp. 319 ◽  
Author(s):  
Xuankun Li ◽  
David K. Yeates
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Maximum Parsimony ◽  
Morphological Characters ◽  
Junior Synonym ◽  
Maximum Parsimony Tree ◽  
New Combinations ◽  
New Genera ◽  
Parsimony Tree

Bombyliinae, one of the largest subfamilies of Bombyliidae, including 78 genera assigned to four tribes, is distributed on all continents except Antarctica. Their larvae are parasitoids, and adults are important pollinators. The Australian Bombyliinae currently has 17 described genera and 87 described species. The purpose of this study is to establish the phylogeny of the Australian members of the subfamily Bombyliinae, including the boundaries and relationships of the genera, using morphological characters. One maximum parsimony tree was generated from 83 morphological characters scored for 50 species, representing all recognised genera and some unplaced species that we considered may represent new genera. Phylogenetic analysis recovered the relationships between the recognised Australian genera of Bombyliinae and clarified generic limits. The group comprises five main clades, with the genus Pilosia being sister to the remaining Bombyliinae. On the basis of this analysis, four new genera are described: Lambkinomyia Li & Yeates, gen. nov., Dissodesma Bowden & Li, gen. nov., Nigromyia Li & Yeates, gen. nov. and Robertsmyia Li & Yeates, gen. nov. Ten new species are described: Dissodesma immaculata Li & Yeates, sp. nov., Dissodesma flava Li & Yeates, sp. nov., Dissodesma smarti Li & Yeates, sp. nov., Nigromyia collessi Li & Yeates, sp. nov., Nigromyia crocea Li & Yeates, sp. nov., Nigromyia flavimana Li & Yeates, sp. nov., Nigromyia longistriata Li & Yeates, sp. nov., Nigromyia pantherina Li & Yeates, sp. nov., Nigromyia parva Li & Yeates, sp. nov. and Nigromyia tomentosa Li & Yeates, sp. nov. Two new junior synonyms are identified: Apiformyia is a junior synonym of Eristalopsis and Brychosoma is a junior synonym of Staurostichus. The following new combinations are proposed: from Apiformyia: Eristalopsis australis (Yeates), comb. nov.; from Bombylius: Dissodesma primogenita (Walker) comb. nov.; from Brychosoma: Staurostichus aureolatus (Walker), comb. nov., Staurostichus hilaris (Walker), comb. nov. and Staurostichus pictipennis (Macquart), comb. nov.; from Laurella: Robertsmyia pallidoventris (Roberts), comb. nov.; from Mandella: Lambkinomyia cinctiventris (Roberts), comb. nov., Lambkinomyia flavovillosa (Roberts), comb. nov., Lambkinomyia pallida (Roberts), comb. nov. and Lambkinomyia rubida (Roberts), comb. nov.; and from Meomyia: Mandella australis (Guérin-Méneville), comb. nov. http://zoobank.org/urn:lsid:zoobank.org:pub:E4A86A19-4AC3-498C-8BDF-B8B501E5A248

Corrigendum to: Morphological phylogeny of the Australian genera of the bee fly subfamily Bombyliinae (Diptera : Bombyliidae) with description of four new genera

2018 ◽  
Vol 32 (2) ◽  
pp. 504
Author(s):  
Xuankun Li ◽  
David K. Yeates
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Maximum Parsimony ◽  
Morphological Characters ◽  
Junior Synonym ◽  
Maximum Parsimony Tree ◽  
New Combinations ◽  
New Genera ◽  
Link Type ◽  
Parsimony Tree

Bombyliinae, one of the largest subfamilies of Bombyliidae, including 78 genera assigned to four tribes, is distributed on all continents except Antarctica. Their larvae are parasitoids, and adults are important pollinators. The Australian Bombyliinae currently has 17 described genera and 87 described species. The purpose of this study is to establish the phylogeny of the Australian members of the subfamily Bombyliinae, including the boundaries and relationships of the genera, using morphological characters. One maximum parsimony tree was generated from 83 morphological characters scored for 50 species, representing all recognised genera and some unplaced species that we considered may represent new genera. Phylogenetic analysis recovered the relationships between the recognised Australian genera of Bombyliinae and clarified generic limits. The group comprises five main clades, with the genus Pilosia being sister to the remaining Bombyliinae. On the basis of this analysis, four new genera are described: Lambkinomyia Li & Yeates, gen. nov., Dissodesma Bowden & Li, gen. nov., Nigromyia Li & Yeates, gen. nov. and Robertsmyia Li & Yeates, gen. nov. Ten new species are described: Dissodesma immaculata Li & Yeates, sp. nov., Dissodesma flava Li & Yeates, sp. nov., Dissodesma smarti Li & Yeates, sp. nov., Nigromyia collessi Li & Yeates, sp. nov., Nigromyia crocea Li & Yeates, sp. nov., Nigromyia flavimana Li & Yeates, sp. nov., Nigromyia longistriata Li & Yeates, sp. nov., Nigromyia pantherina Li & Yeates, sp. nov., Nigromyia parva Li & Yeates, sp. nov. and Nigromyia tomentosa Li & Yeates, sp. nov. Two new junior synonyms are identified: Apiformyia is a junior synonym of Eristalopsis and Brychosoma is a junior synonym of Staurostichus. The following new combinations are proposed: from Apiformyia: Eristalopsis australis (Yeates), comb. nov.; from Bombylius: Dissodesma primogenita (Walker) comb. nov.; from Brychosoma: Staurostichus aureolatus (Walker), comb. nov., Staurostichus hilaris (Walker), comb. nov. and Staurostichus pictipennis (Macquart), comb. nov.; from Laurella: Robertsmyia pallidoventris (Roberts), comb. nov.; from Mandella: Lambkinomyia cinctiventris (Roberts), comb. nov., Lambkinomyia flavovillosa (Roberts), comb. nov., Lambkinomyia pallida (Roberts), comb. nov. and Lambkinomyia rubida (Roberts), comb. nov.; and from Meomyia: Mandella australis (Guérin-Méneville), comb. nov. <a ext-link-type=

scholarly journals Systematics and phylogeny of the Neotropical treehopper subfamily Nicomiinae (Hemiptera, Membracidae)

2005 ◽  
Vol 22 (1) ◽  
pp. 231-283 ◽  
Author(s):  
Jesse L. Albertson ◽  
Christopher H. Dietrich
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Morphological Characters ◽  
Junior Synonym ◽  
Type Material ◽  
New Combinations ◽  
Primary Type ◽  
New Synonymy

Morphological characters of adults of the treehopper subfamily Nicomiinae Haupt, 1929 (Hemiptera, Membracidae) including seven genera (Eudonica gen. nov.; Euwalkeria Goding, 1926; Holdgatiella Evans, 1962; Nicomia Stål, 1858; Nodonica Dietrich, McKamey& Deitz, 2001; Stalomia gen. nov.; and Tolania Stål, 1858) and 22 species (16 new) are described and illustrated. Keys are provided for genera and for species of Euwalkeria, Holdgatiella, and Nicomia. Nomenclatural changes, based on study of the primary type material of 15 species, include three new combinations, one new synonymy, and reinstatement of one junior synonym. Eudonica has one species, Eudonica nanella sp. nov.; Euwalkeria has five species, including four new species: E. colorata sp. nov., E. distincta sp. nov., E. perdita sp. nov., E. rubrica sp. nov.; Holdgatiella has two species, one of which is described as new: Holdgatiella chiloensis sp. nov.; Nicomia has twelve species, nine of which are described as new: N. buccina sp. nov., N. harenosa sp. nov., N. inscripta sp. nov., N. jucunda sp. nov., N. monticola sp. nov., N. nigrifasciata sp. nov., N. notidana sp. nov., N. pulchella sp. nov., N. serrata sp. nov.; Nodonica has one species, Nodonica bispinigera Dietrich, McKamey & Deitz; and Stalomia has one species, Stalomia veruta sp. nov. Tolania contains eleven previously described species and nearly 60 new species, which will be treated in a later publication. Three new combinations are proposed: one species described in Nicomia is placed in the tribe Abelini (Centrotinae) as Abelus retrospinosus (Lethierry) comb. nov.; one species previously placed in Nicomia is transferred to the genus Tolania as T. obliqua (Walker, 1858), comb. nov.; one species described in Holdgatiella is placed in the genus Tolania as T. stria (Cryan & Deitz, 2002), comb. nov. One new synonymy is proposed: Hoplophera [sic] cicadoides Walker, 1862, syn. nov., a junior synonym of Nicomia interrupta Stål, 1858. Nicomia subfasciata Stål, 1858, is reinstated. The results of a phylogenetic analysis based on morphology are presented, illustrating the hypothesized relationships among species and genera of Nicomiinae. The analysis supports the broader concept of Nicomiinae proposed here. The monophyly of this group is supported by eleven characters, and all of the included genera are also monophyletic.

scholarly journals Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species

ZooKeys ◽  
2020 ◽  
Vol 985 ◽  
pp. 71-126
Author(s):  
Jose I. Martinez
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Morphological Characters ◽  
The Other ◽  
South American ◽  
New Combinations ◽  
New Genera ◽  
The South ◽  
Molecular Analyses ◽  
One New Species

The endemic Neotropical genus Gaujonia Dognin is revised. Morphological characters and a phylogenetic analysis demonstrate paraphyletic relationships among the species. Four different groups are interpreted to represent four different genera. The G. arbosi group is the only remaining clade in the genus Gaujonia, and the other groups have been arranged into three new genera: Milleranagen. nov., Oculicattusgen. nov., and Cicadoformagen. nov. Additionally, two other genera Cicadomorphusgen. nov., and Gaujopteragen. nov. were found using morphological and molecular analyses based on some specimens that were misidentified as Gaujonia spp. A total of five new genera, three new combinations (Cicadoforma vau-nigrum Hampson, comb. nov., Oculicattus renifera Hampson, comb. nov., and Millerana arbosioides Dognin, comb. nov.) and 21 new species (Cicadoforma ocelotussp. nov., Cicadomorphus chicharrasp. nov., Cicadomorphus chuyasp. nov., Cicadomorphus falkasiskasp. nov., Cicadomorphus lilianaesp. nov., Gaujonia bichusp. nov., Gaujonia chiqyaqsp. nov., Gaujonia kanakusikasp. nov., Gaujonia sourakovisp. nov., Gaujoptera amsasp. nov., Millerana austinisp. nov., Millerana cajassp. nov., Millerana cundinamarquensissp. nov., Millerana matthewsaesp. nov., Millerana tigrinasp. nov., Oculicattus bolivianasp. nov., Oculicattus brehmisp. nov., Oculicattus incasp. nov., Oculicattus raizaesp. nov., Oculicattus schmidtisp. nov., and Oculicattus uturunkusp. nov.) are established.

Revision and phylogenetic analysis of the Papuan weevil genus Thyestetha Pascoe (Coleoptera, Curculionidae, Cryptorhynchinae)

Zootaxa ◽  
2012 ◽  
Vol 3355 (1) ◽  
pp. 1 ◽  
Author(s):  
JONAS EBERLE ◽  
RENE TÄNZLER ◽  
ALEXANDER RIEDEL
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Maximum Parsimony ◽  
Phylogenetic Analyses ◽  
New Guinea ◽  
Morphological Characters ◽  
Junior Synonym ◽  
Maximum Parsimony Analysis ◽  
Parsimony Analysis

Thyestetha Pascoe, an apterous genus of Papuan Cryptorhynchinae is revised. It contains two previously describedspecies, T. carbonaria Faust and T. nitida Pascoe, plus seven new species: T. discolor sp. n., T. emarginata sp. n.,T. glabra sp. n., T. lubrica sp. n., T. oblita sp. n., T. puncticollis sp. n., and T. splendens sp. n.. Lectotypes are designatedfor T. carbonaria, T. nitida, and Pantoxistus rubripennis Chevrolat which is a junior synonym of T. nitida (syn. n.). Thegenus and its species are described and the characters relevant for their identification are illustrated. A key to the speciesis provided. Phylogenetic analyses are performed, utilizing molecular and morphological approaches. A maximumlikelihood tree including 5 Thyestetha species and 2 outgroups is presented, based on 3 mitochondrial and 4 nuclearmarkers. A maximum parsimony analysis of 19 morphological characters includes all described species. Thyestetha ispresumably closely related to Telaugia Pascoe and Elichora Pascoe. The group is restricted to New Guinea and some smaller neighboring islands.

Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae)

Zootaxa ◽  
2020 ◽  
Vol 4796 (1) ◽  
pp. 1-322
Author(s):  
RENATO JOSE PIRES MACHADO ◽  
JOHN DAVID OSWALD
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Phylogenetic Relationships ◽  
Taxonomic Revision ◽  
New Guinea ◽  
Morphological Characters ◽  
Valid Species ◽  
New Combinations ◽  
New Genera ◽  
Parsimony Analysis

The species of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae) are revised. Prior to this work Periclystina comprised 10 genera and 63 species endemic to Australia (62 spp.) and New Guinea (1 sp.). In this work two former genera are synonymized and four new genera are proposed (for a total of 12 valid genera), and six former species are synonymized and 16 new species are proposed (for a total of 73 valid species). A parsimony analysis utilizing 62 morphological characters is used to infer phylogenetic relationships among all 73 species, and suitable outgroups. Based on the results of the phylogenetic analysis, the tribal and/or generic assignments of 55 (75%) of former Periclystina species are modified here. Periclystina and its two largest genera—Austrogymnocnemia and Glenoleon—were all recovered as polyphyletic in their former circumscriptions. Ten species—placed here in six genera: Anomaloplectron, Csiroleon, Fissuleon gen. nov., Franzenia, Fusoleon and Latileon gen. nov.—clustered phylogenetically with Acanthoplectron and are transferred to the tribe Acanthoplectrini. The remaining 63 species—placed in six additional genera: Austrogymnocnemia, Glenoleon, Megagonoleon gen. nov., Minyleon gen. nov., Periclystus and Riekoleon—form a monophyletic Periclystus genus group and are transferred to the tribe Dendroleontini. All 73 species are (re)described and illustrated. New identification keys are provided for the tribes of Dendroleontinae, for the Australian genera of Acanthoplectrini and Dendroleontini, and for each of the non-monotypic genera treated. In addition, five new lectotypes are designated and 35 new combinations are created. 

A taxonomic review and phylogeny of the genera Hoplopisthius Senna, 1892 and Carcinopisthius Kolbe, 1892 (Coleoptera: Brentidae, Hoplopisthiini)

Zootaxa ◽  
2010 ◽  
Vol 2516 (1) ◽  
pp. 1 ◽  
Author(s):  
ANTOINE MANTILLERI
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Maximum Parsimony ◽  
New Guinea ◽  
Morphological Characters ◽  
Identification Key ◽  
Generic Level ◽  
Australian Species ◽  
Taxonomic Review ◽  
One New Species

The genera Hoplopisthius and Carcinopisthius are reviewed. All the species are redescribed and illustrated and an updated identification key and distributional maps are provided. One new species from New Guinea is described (Hoplopisthius maximus n. sp.) and two new synonymies are proposed: Hoplopisthius celebensis Kolbe, 1892 = H. trichemerus Senna, 1892, n. syn. and Carcinopisthius lamingtoni Damoiseau, 1987 = C. forcipitiger Damoiseau, 1987, n. syn. Phylogenetic analysis using PAUP (maximum parsimony) was performed using 25 morphological characters of adults. This analysis shows the group Hoplopisthius + Carcinopisthius is monophyletic, but Carcinopisthius alone is paraphyletic. Nomenclatural changes at the generic level are made to reconcile nomenclature and phylogeny: Hoplopisthius is preserved; Carcinopisthius is downgraded to the rank of subgenus for the two oriental species H. oberthueri and H. fruhstorferi; and Pseudotaphroderes is resurrected as a third subgenus and includes all New-Guinean and Australian species.

Revision of the Nepalese genus Microplinthus Zherichin, 1987 (Coleoptera: Curculionidae: Molytinae), with description of 25 new species

Zootaxa ◽  
2020 ◽  
Vol 4794 (1) ◽  
pp. 1-63
Author(s):  
MASSIMO MEREGALLI
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Bayesian Inference ◽  
Maximum Parsimony ◽  
Morphological Characters ◽  
Type Locality ◽  
Central Nepal

The Nepalese genus Microplinthus Zherichin, 1987 is revised. The following 25 new species are described (type locality in parentheses): M. annapurnae (Annapurna Himal, Rambrong Danda, 28°23’N 84°13’E), M. arunensis (Khandbari Distr., forest S. Mansimgma, 27°30’N 87°20’E), M. brevipennis (Manaslu Himal, Bhara Pokhari, 28°18’N 84°28’E), M. durga (W Pokhara, near Ghorepani, 28°24’N 83°42’E), M. franzi (Kaligandakhi valley, between Lethe and Tukuche, 28°36’N 83°38’E), M. ganesha (Ganesh Himal, Rupchet, 28°05’N 85°09’E), M. godawaricus (Kathmandu, Phulchowki, 27°34’N 85°24’E), M. gupta (Helambu Himal, upper Chipling, 27°53’N 85°28’E), M. helambuensis (Central Nepal, between Mulkharka and Thare Pati, 28°08’N 85°22’E–28°02’N 85°29’E), M. jaegeri (W Pokhara, Mt. Panchase, 28°13’N 83°48’E), M. karnalicus (Karnali Distr. 5 km E Charta, 29°18’N 82°20’E), M. kaskianus (Kaski Distr., above Pothana, 28°19’N 83°51’E), M. kaumarya (Karnali Distr., Gotichaur valley, 29°12’10”N 82°18’56”E), M. khandbariensis (Khandbari Distr., above Seduwa, 27°34’N 87°15’E), M. koshianus (Koshi Prov., SW Chichilla, 27°27.01’N 87°13.13’E), M. letheensis (Kaligandakhi valley, between Lethe and Tukuche, 28°36’N 83°38’E), M. longipennis (Manaslu Himal, Bhara Pokhari lekh, 28°18’24”N 84°28’06”E), M. messneri (Panchtar Distr., Dhorpar Kharka, 27°05’N 87°55’E), M. newarorum (Phulchowki Distr., Lalitpur, 27°34’N 85°24’E), M. phulchowkianus (Kathmandu, Phulchowki, 27°34’N 85°24’E), M. rugosus (Solokhumbu Distr., Lamiura Danda, 27°34’N 86° 30’E), M. schmidti (Manaslu Himal, Dudh Pokhari lekh, 28°20’N 84°35’E), M. sherpa (Khandbari Distr., Bakan, 27°38’N 87°12’E), M. tamanus (Rolwaling Himal, upper Simigau, 27°51’N 86°15’E), M. yeti (Hinku Distr., Drangka Khola bridge, 27°35’N 86°47’E). A key for the identification of all species of Microplinthus is given. A phylogenetic analysis based on morphological characters of the species referable to the genera Falsanchonus, Microniphades and Microplinthus was carried out, implementing Bayesian Inference and Maximum Parsimony. Results indicate that the genera Falsanchonus and Microniphades are paraphyletic and are here treated as junior synonyms of Microplinthus. 

Revision of Asytesta Pascoe, 1865, with comments on the phylogeny of the Indo-Australian crowned weevils (Coleoptera: Curculionidae: Cryptorhynchinae)

Zootaxa ◽  
2012 ◽  
Vol 3462 (1) ◽  
pp. 1 ◽  
Author(s):  
GREGORY P. SETLIFF
Keyword(s):  
New Species ◽  
Morphological Characters ◽  
Junior Synonym ◽  
New Combinations ◽  
Monotypic Genus ◽  
Species Status ◽  
Distribution Maps ◽  
Line Drawings ◽  
New Synonyms ◽  
Species Groups

The Indo-Australian crowned weevil genus Asytesta Pascoe, 1865 is revised. Forty-one species are recognized, including18 that are new: A. alexandriae, A. alexriedeli, A. allisoni, A. biakana, A. cheesmanae, A. concolora, A. emarginata, A.fayae, A. frontalis, A. gressitti, A. julieae, A. marginalis, A. morobeana, A. sedlaceki, A. thompsoni, A. tuberculata, A.vivienae, and A. woodlarkiana, new species. One subspecies, A. lugubris bidentata Voss is elevated to species status, A.bidentata Voss, new status. Four species are newly synonymized: A. circulifera Lea, 1928 = A. rata Heller, 1910, A.definita Faust, 1898 = A. humeralis Pascoe, 1865, A. granulifera Lea, 1928 = A. aucta Faust, 1898, and A. setipes Lea,1928 = A. lugubris Heller, 1895 new synonyms. Six new species groups are proposed. Lectotypes are designated for 18species. Two species are transferred from Asytesta to other genera: A. maura Pascoe to Microporopterus Lea and A.ypsilon Heller to Meroleptus Faust, new combinations. A checklist and key for all crowned weevil genera, key to speciesgroups and species of Asytesta, adult habitus illustrations, distribution maps, and line drawings of diagnostic charactersare provided.A phylogeny for the genus based on 82 adult morphological characters (187 states) for 41 ingroup taxa is alsopresented. All genera and species of the crowned weevil group as redefined here (including Cyamomistus Heller,Eudyasmus Pascoe, Glochinorhinus Waterhouse, Nothotragopus Zimmerman, Panopides Pascoe, and Zygara Pascoe),were included in the analysis to test the monophyly of Asytesta. Monophyly of Asytesta was supported only with thesynonymy of the monotypic genus Zygara. Accordingly, Zygara is a new junior synonym of Asytesta and Zygara doriae (Kirsch) is returned to its original combination with Asytesta; A. doriae Kirsch resurrected status.

Phylogenetic analysis of Lumbrineridae Schmarda, 1861 (Annelida: Polychaeta)

Zootaxa ◽  
2006 ◽  
Vol 1332 (1) ◽  
pp. 1 ◽  
Author(s):  
LUIS F. CARRERA-PARRA
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Type Species ◽  
Phylogenetic Reconstruction ◽  
Morphological Characters ◽  
New Genera ◽  
The Family ◽  
Three New Species

This study represents the first phylogenetic reconstruction of lumbrinerid genera using parsimony analyses of 38 morphological characters. Following higher-level phylogenetic analysis, Oenone (Oenonidae) was selected as outgroup. The analysis was restricted to type species for each genus, yielded 24 equally parsimonious trees, which after successive weighting were reduced to one tree (CI= 0.7396). The topology of this tree revealed the separation of the family into four main clades: 1. Lysarete, 2. Arabellonereis, 3. Scoletoma, Lumbrineris, Hilbigneris gen. nov., Kuwaita, Lumbricalus, Sergioneris gen. nov. and Eranno, and 4. Abyssoninoe, Cenogenus, Lumbrinerides, Lumbrineriopsis, Augeneria, Loboneris gen. nov., Gallardoneris gen. nov., Helmutneris gen. nov., and Gesaneris gen. nov.; the position of Ninoe is unclear. A diagnosis of each genus is provided, including the description of six new genera and three new species. A key to lumbrinerid genera is included.

Taxonomic review of the Craniophora s. l. (Lepidoptera, Noctuidae, Acronictinae) generic complex with description of 8 new genera and 13 new species

Zootaxa ◽  
2017 ◽  
Vol 4355 (1) ◽  
pp. 1
Author(s):  
ÁDÁM KISS
Keyword(s):  
New Species ◽  
Morphological Characters ◽  
New Combinations ◽  
New Genera ◽  
Genital Morphology ◽  
Male And Female ◽  
Taxonomic Review ◽  
New States ◽  
First Time ◽  
Lepidoptera Noctuidae

Present paper contains the description of 8 new genera, separated from Craniophora (Harmandicrania, Graesericrania, Eurypterocrania, Turnerinycta, Fascionycta, Berionycta, Draudtinycta and Sinonycta gen. n.) and 13 new species (Harmandicrania barnandi, H. tathabayandi, H. brunneocinerea, H. sinoandi, H. peninsularis, H. nipponica, Berionycta limbata, B. ponticamima, B. nigra, B. orbicularis, B. behouneki, B. berioi and B. beckroberti spp. n.). Based on external and genital morphology and characters of the abdominal segments the taxonomic context of the genus Craniophora was reduced, the genus Megalonycta was extended. 22 new combinations and 11 new states were introduced. 8 lectotypes and 4 neotypes were designated. With 6 plates of morphological characters and 231 figures in 33 plates. The male and female last abdominal segments of most of the species are figured for the first time. 

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