A systematic and phylogenetic revision of the family Elipsocidae (Insecta : Psocoptera), with the erection of two new families: Lesneiidae and Sabulopsocidae

2004 ◽  
Vol 18 (2) ◽  
pp. 157 ◽  
Author(s):  
T. R. New ◽  
E. R. Schmidt

The phylogeny of the Elipsocidae is reappraised to aid understanding of generic relationships within the family, leading to its clear definition as a monophyletic entity and clarification of its position within the infraorder Homilopsocidea. Examination of new material and generic-level revision results in descriptions of ten new elipsocid genera: Prionotodrilus, gen. nov. for P. parvus (Smithers & Thornton), comb. nov. and P. serratus (Schmidt & Thornton), comb. nov.; Ausysium, gen. nov. for A. joyceorum, sp. nov.; Diademadrilus, gen. nov. for D. annulatus (Smithers), comb. nov., for which a description of the female is given, and D. masseyi (New), comb. nov.; Euryphallus, gen. nov. for E. badonneli (New & Thornton), comb. nov., E. cinqueportsae (Thornton & New), comb. nov., E. cooki (Thornton & New), comb. nov., E. defoei (Thornton & New), comb. nov., E. selkirki (Thornton & New), comb. nov., E. skottsbergi (Thornton & New), comb. nov. and E. stigmaticus (Tillyard), comb. nov.; Gondwanapsocus, gen. nov. for G. australis, sp. nov.; Onychophallus, gen. nov. for O. diemenensis, sp. nov.; Psocophloea, gen. nov. for P. sarahae, sp. nov.; Telmopsocus, gen. nov. for T. waldheimensis, sp. nov.; Villopsocus, gen. nov. for V. tasmaniensis, sp. nov., and Weddellopsocus, gen. nov. for W. avius (Smithers), comb. nov., W. carrilloi (Thornton & Lyall), comb. nov., W. flavus (Thornton & Lyall), comb. nov., W. griseus (New & Thornton), comb. nov. and W. valdiviensis (Blanchard), comb. nov. A rediagnosis of Spilopsocus is given, the male of Sabulopsocus tractuosus Smithers is described and Nepiomorpha phragmitella Smithers is transferred to Clinopsocus New. We provide a phylogenetic analysis to assess the monophyly of Elipsocidae. Sixty-one characters are scored for 38 taxa, including the putative new genera and four outgroups. Monophyly of Elipsocidae is supported with the exclusion of four genera: Palmicola Mockford is transferred to Mesopsocidae; Sabulopsocus Smithers and Moapsocus, gen.�nov. for M. angelicus, sp. nov. are placed in the new family Sabulopsocidae; and Drymopsocus Smithers, which is the sister-taxon to Elipsocidae, is incertae sedis. Other taxa previously regarded as elipsocids are reassigned: Elipsocus impressus (Hagen), Elipsocus modestus Banks and Elipsocus boops (Hagen) are redescribed and respectively placed in Valenzuela Navás (Caeciliusidae), Dasydemella Enderlein (Dasydemellidae) and Mesopsocus Kolbe (Mesopsocidae). Lesneia Badonnel is placed in the new family Lesneiidae. The position of Eolachesilla Badonnel is discussed briefly. A key to genera and comment on characters diagnosing the family Elipsocidae is presented. We retain two subfamilies and discuss the merits of our proposed higher classification. A brief biogeographical interpretation suggests that the family has a southern origin and may be as old as the Mesozoic.

2000 ◽  
Vol 69 (4) ◽  
pp. 223-250 ◽  
Author(s):  
Carrie E. Schweitzer ◽  
Rodney M. Feldmann

New fossils referable to the Cancridae Latreille, 1802 extend the known stratigraphic range of the family into the middle Eocene and the geographic range into South America. Each genus within the family has been reevaluated within the context of the new material. A suite of diagnostic characters for each cancrid genus makes it possible to assign both extant and fossil specimens to genera and the two cancrid subfamilies, the Cancrinae Latreille, 1802, and Lobocarcininae Beurlen, 1930, based solely upon dorsal carapace morphology. Cheliped morphology is useful in assigning genera to the family but is significantly less useful at the subfamily and generic level. Each of the four subgenera sensu Nations (1975), Cancer Linnaeus, 1758, Glebocarcinus Nations, 1975, Metacarcinus A. Milne Edwards, 1862, and Romaleon Gistl, 1848, are elevated to full generic status. Additionally, three new genera and three new species accommodate the new, as well as some previously described taxa, and include Anatolikos new genus, Anisospinos berglundi new genus and species, and Notocarcinus sulcatus new genus and species and several new combinations. Recognition of new genera and reassignment of several species within the Cancrinae indicates that that subfamily may have arisen in the southern hemisphere, contrary to the previous interpretation of the subfamily as a primarily North Pacific or Tethyan group. The Lobocarcininae was primarily a Tethyan group.


2011 ◽  
Vol 20 (1) ◽  
pp. 161-173
Author(s):  
A.P. Kassatkina

Resuming published and own data, a revision of classification of Chaetognatha is presented. The family Sagittidae Claus & Grobben, 1905 is given a rank of subclass, Sagittiones, characterised, in particular, by the presence of two pairs of sac-like gelatinous structures or two pairs of fins. Besides the order Aphragmophora Tokioka, 1965, it contains the new order Biphragmosagittiformes ord. nov., which is a unique group of Chaetognatha with an unusual combination of morphological characters: the transverse muscles present in both the trunk and the tail sections of the body; the seminal vesicles simple, without internal complex compartments; the presence of two pairs of lateral fins. The only family assigned to the new order, Biphragmosagittidae fam. nov., contains two genera. Diagnoses of the two new genera, Biphragmosagitta gen. nov. (type species B. tarasovi sp. nov. and B. angusticephala sp. nov.) and Biphragmofastigata gen. nov. (type species B. fastigata sp. nov.), detailed descriptions and pictures of the three new species are presented.


2003 ◽  
Vol 60 (3) ◽  
pp. 533-568 ◽  
Author(s):  
J. C. MANNING ◽  
P. GOLDBLATT ◽  
M. F. FAY

A revised generic synopsis of sub-Saharan Hyacinthaceae is presented, based on a molecular phylogenetic analysis of the family. Generic rank is accorded only to reciprocally monophyletic clades that can be distinguished by recognizable morphological discontinuities, thereby permitting an appropriate generic assignment of species not included in the analysis. Three subfamilies are recognized within the region. Subfamily Ornithogaloideae, characterized by flattened or angular seeds with tightly adhering testa, is considered to include the single genus Ornithogalum, which is expanded to include the genera Albuca, Dipcadi, Galtonia, Neopatersonia and Pseudogaltonia. Recognizing any of these segregates at generic level renders the genus Ornithogalum polyphyletic, while subdivision of Ornithogalum into smaller, morphologically distinguishable segregates in order to preserve the monophyly of each is not possible. Subfamily Urgineoideae, characterized by flattened or winged seeds with brittle, loosely adhering testa, comprises the two mainland African genera Bowiea and Drimia. The latter is well circumscribed by its deciduous, short-lived perianth and includes the previously recognized genera Litanthus, Rhadamanthus, Schizobasis and Tenicroa. The monotypic Madagascan Igidia is provisionally included in the subfamily as a third genus on the basis of its seeds, pending molecular confirmation of its relationships. Subfamily Hyacinthoideae resolves into three clades, distinguished as tribes Hyacintheae (strictly northern hemisphere and not treated further), Massonieae and Pseudoprospereae tribus nov. Full descriptions and a key to their identification are provided for all genera. New combinations reflecting the generic circumscriptions adopted here are made for most African and all Indian and Madagascan species.


2021 ◽  
pp. SP521-2020-249
Author(s):  
Daran Zheng ◽  
Edmund A. Jarzembowski ◽  
De Zhuo ◽  
André Nel

AbstractHemiphlebiidae are the most basal lestomorphan family following the latest phylogenetic analysis of the Zygoptera: this unique damselfly family today contains one relict species found in the wetlands of Australia. It was, however, very diverse and widespread during the Mesozoic. Nevertheless, very few species were known obscuring the origination and early evolution of the family. Here we propose a new stem hemiphlebioid taxon (Protohemiphlebiidae Zheng, Jarzembowski & Nel, fam. nov.) based on a new genus and two species: Protohemiphlebia zhangi Zheng, Jarzembowski & Nel, sp. nov. and Protohemiphlebia meiyingae Zheng, Jarzembowski & Nel, sp. nov. The new family shares the characters of both Hemiphlebiidae and Coenagrionoidea, but it is more closely related to Hemiphlebiidae in having the pterostigma with a ‘star-shaped’ microsculpture, and AA originating from the wing base slightly distal of Ax0. Protohemiphlebia Zheng, Jarzembowski & Nel, gen. nov. is further considered to belong to the stem group of Hemiphlebioidea, instead of belonging to the Hemiphlebiidae, in possessing pretibial combs and a weakly kinked RP1 below the Pt-brace. The new damselflies will help to calibrate the origin of Hemiphlebiidae, which could be earlier than their current oldest records in the Kimmeridgean (Late Jurassic).


2010 ◽  
Vol 24 (6) ◽  
pp. 573 ◽  
Author(s):  
Guillermo Kuschel ◽  
Richard A. B. Leschen

An overview of the taxa of Rhinorhynchinae (Nemonychidae) is presented. A phylogenetic analysis of the 19 extant genera of the Rhinorhynchinae and one fossil genus from the Lower Cretaceous (Cratomacer Zherikhin & Gratshev), based on 29 characters of adults, larvae and host plant associations, shows three monophyletic groups, treated as tribes (Rhinorhynchini, Mecomacerini and Rhynchitomacerini). Primitive associations are unknown for Rhinorhynchinae and early host use diversification included associations with Ranunculaceae (Nemonyx Redtenbacher), Pinaceae (Cimberis Gozis), Nothofagaceae (Rhynchitomacerini) and Podocarpaceae (Rhinorhynchini). While Mecomacerini diversified on Araucariaceae, within Rhinorhynchini there was a single reversal to Araucariaceae and a shift in Atopomacer Kuschel to Pinaceae. Placement of Cratomacer into Mecomacerini is consistent with geological and molecular data that suggest gymnosperms may have been the primitive hosts for the family. Three new genera are described in Rhinorhynchinae: Araucomacer, gen. nov. (type species A. hirticeps Kuschel) from Chile, Idiomacer, gen. nov. (type species I. basicornis, sp. nov.) from New Caledonia, and Zimmiellus, gen. nov. (type species Z. fronto, sp. nov.) from Queensland, Australia, while Stenomacer Kuschel, stat. nov. from Chile is reinstated from synonymy. Seven species are described as new: Atopomacer grandifurca, sp. nov. from Costa Rica and Panama, A. obrieni, sp. nov. from Mexico, A. panamensis, sp. nov. from Panama, A. pini, sp. nov. from Mexico, A. podocarpi, sp. nov. from Venezuela, Basiliogeus dracrycarpi, sp. nov. from western New Guinea and B. inops, sp. nov. from Queensland. Rhynchitomacer rufus Kuschel is a new junior subjective synonym of R. nigritus Kuschel, syn. nov., R. viridulus Kuschel is a new junior subjective synonym of R. flavus Voss, syn. nov. and Stenomacer fuscus Kuschel is a new junior subjective synonym of S. vernus Kuschel, syn. nov. Keys to genera of Rhinorhynchinae and to species of Atopomacer, Rhynchitomacer and Stenomacer are included. Diagnoses of the three extant subfamilies of Nemonychidae are included.


2019 ◽  
Vol 33 (6) ◽  
pp. 892
Author(s):  
Adriano B. Kury ◽  
Abel Pérez-González ◽  
Daniel N. Proud

The laniatorean family Phalangodidae has been largely reduced to a core of Holarctic species; however, many taxa were never formally transferred to other families. We examined a group of harvestmen related to Beloniscus Thorell, 1891, from South-east Asia and determined the nature of the relationships of the ‘Beloniscus-like harvestmen’, herein described as the new family Beloniscidae, fam. nov., in the broad context of Laniatores. Based on a molecular phylogenetic analysis of a broad representative laniatorean Sanger-sequences dataset we found support for our taxonomic hypotheses that (1) inclusion of Beloniscidae, fam. nov. in Phalangodidae would render the family non-monophyletic, and (2) Beloniscidae, fam. nov. represents a lineage that is morphologically and genetically distinct from all other known laniatorean families. The new family Beloniscidae is endemic to South-east Asia and comprises 37 species in two new subfamilies: Beloniscinae, subfam. nov. and Buparinae, subfam. nov. Beloniscinae includes the genera Beloniscellus Roewer, 1931, Beloniscops Roewer, 1949, Belonisculus Roewer, 1923, Beloniscus (type genus) and Kendengus Roewer, 1949. Buparinae includes the genera Buparellus Roewer, 1949, Bupares Thorell, 1889 (type genus) and Buparomma Roewer, 1949. Members of Beloniscidae are relatively homogeneous in their external morphology, but are recognised by well marked genitalic features. Male genitalia are described for exemplar species, and the morphology is compared with that of Epedanidae. Names are given to two new genitalic structures: sella curulis and pseudocalyx. http://zoobank.org/urn:lsid:zoobank.org:act:540BCE49-6F2E-4372-BFD4-D3C6068F045D http://zoobank.org/urn:lsid:zoobank.org:act:FE8B011B-AA6D-4E6C-A321-123542A0563F http://zoobank.org/urn:lsid:zoobank.org:act:E4A559CD-170A-40F2-924A-541C22974344


2017 ◽  
Vol 31 (5) ◽  
pp. 566 ◽  
Author(s):  
Michael G. Rix ◽  
Robert J. Raven ◽  
Barbara Y. Main ◽  
Sophie E. Harrison ◽  
Andrew D. Austin ◽  
...  

The Australasian spiny trapdoor spiders of the family Idiopidae (subfamily Arbanitinae) are revised at the generic level, using a multi-locus molecular phylogenetic foundation and comprehensive sampling of all known lineages. We propose a new family- and genus-group classification for the monophyletic Australasian fauna, and recognise 10 genera in four tribes. The Arbanitini Simon includes Arbanitis L. Koch, 1874 (61 species), Blakistonia Hogg, 1902 (one species) and Cantuaria Hogg, 1902 (43 species). The Aganippini Simon includes Bungulla Rix, Main, Raven & Harvey, gen. nov. (two species), Eucanippe Rix, Main, Raven & Harvey, gen. nov. (one species), Eucyrtops Pocock, 1897 (two species), Gaius Rainbow, 1914 (one species) and Idiosoma Ausserer, 1871 (14 species). The Cataxiini Rainbow and Euoplini Rainbow include just Cataxia Rainbow, 1914 (11 species) and Euoplos Rainbow, 1914 (12 species), respectively. Two distinctive new genera of Aganippini are described from Western Australia, and several previously valid genera are recognised as junior synonyms of existing genus-group names, including Misgolas Karsch, 1878 (= Arbanitis; new synonymy), Aganippe O. P.-Cambridge, 1877 (= Idiosoma; new synonymy) and Anidiops Pocock, 1897 (= Idiosoma; new synonymy). Gaius stat. rev. is further removed from synonymy of Anidiops. Other previously hypothesised generic synonyms are supported by both morphology and molecular phylogenetic data from 12 genes, including the synonymy of Neohomogona Main, 1985 and Homogona Rainbow, 1914 with Cataxia, and the synonymy of Albaniana Rainbow & Pulleine, 1918, Armadalia Rainbow & Pulleine, 1918, Bancroftiana Rainbow & Pulleine, 1918 and Tambouriniana Rainbow & Pulleine, 1918 with Euoplos. At the species level, the identifications of Eucy. latior (O. P.-Cambridge, 1877) and I. manstridgei (Pocock, 1897) are clarified, and three new species are described: Bungulla bertmaini Rix, Main, Raven & Harvey, sp. nov., Eucanippe bifida Rix, Main, Raven & Harvey, sp. nov. and Idiosoma galeosomoides Rix, Main, Raven & Harvey, sp. nov., the latter remarkable for its phragmotic abdominal morphology. The Tasmanian species Mygale annulipes C. L. Koch, 1842 is here transferred to the genus Stanwellia Rainbow & Pulleine, 1918 (family Nemesiidae), comb. nov., Arbanitis mestoni Hickman, 1928 is transferred to Cantuaria, comb. nov. and Idiosoma hirsutum Main, 1952 is synonymised with I. sigillatum (O. P.-Cambridge, 1870), new synonymy. In addition to the morphological synopses and an illustrated key to genera, molecular diagnoses are presented for all nominal taxa, along with live habitus and burrow images to assist in field identification. The Australasian idiopid fauna is highly diverse, with numerous new species known from all genera. As a result, this study provides a taxonomic and nomenclatural foundation for future species-level analyses, and a single reference point for the monographic documentation of a remarkable fauna. http://zoobank.org/?lsid=urn:lsid:zoobank.org:pub:BACE065D-1EF9-40C6-9134-AADC9235FAD8


Phytotaxa ◽  
2014 ◽  
Vol 181 (3) ◽  
pp. 151 ◽  
Author(s):  
Yuya Inoue ◽  
Hiromi Tsubota

Based on our molecular phylogenetic analysis of haplolepideous mosses with concatenated sequences of chloroplast rps4 and rbcL genes, a new family Timmiellaceae is erected to accommodate the genera Timmiella and Luisierella, both of which have been formerly included in the family Pottiaceae.  The family Timmiellaceae is resolved as a second-branching clade together with Distichium (Distichiaceae) within the Dicranidae (haplolepideous moss) lineages and phylogenetically distinct from the Pottiaceae.  Reassessment of morphological characters suggests that a combination of the characters: 1) adaxially bulging and abaxially flat leaf surfaces, 2) sinistrorse or straight peristomes, when present, and 3) sinistrorsely arranged operculum cells is unique to Timmiellaceae and discriminates it from other haplolepideous moss families.


2010 ◽  
Vol 8 (3) ◽  
pp. 385-568 ◽  
Author(s):  
Juan Marcos Mirande

The family Characidae is the most diverse among Neotropical fishes. Systematics of this family are mainly based on pre-cladistic papers, and only recently a phylogenetic hypothesis for Characidae was proposed by the author. That phylogeny was based on 360 morphological characters studied for 160 species, including representatives of families related to Characidae. This paper is based on that phylogenetic analysis, with the analyzed characters described herein and documented, accompanied by comparisons of their definition and coding in previous papers. Synapomorphies of each node of the proposed phylogeny are listed, comparisons with previous classifications provided, and autapomorphies of the analyzed species listed. Taxonomic implications of the proposed classification and the position of the incertae sedis genera within Characidae are discussed. A discussion of the phylogenetic information of the characters used in the classical systematics of the Characidae is provided.


1995 ◽  
Vol 69 (4) ◽  
pp. 670-685 ◽  
Author(s):  
D. I. Mackinnon ◽  
T. N. Smirnova

Four species of megathyrid brachiopods from the Lower Cretaceous (Berriasian) of Crimea, Ukraine, and one species of megathyrid from the lower Upper Cretaceous (Cenomanian) of England are reinvestigated using scanning electron microscopy. The species are variously assigned to one existing genus, Praeargyrotheca Smirnova (P. hexaplicata (Smirnova) and P. megatrema (J. de C. Sowerby)) and two new genera, Krimargyrotheca (K. concinna (Smirnova) and K. balkii n. gen. and n. sp.) and Evargyrotheca (E. alta (Smirnova)). Taxa are distinguished principally on differences in shell shape, plication, and, in particular, surface microornament. The shell microstructure of all five species is investigated with particular emphasis on endopunctation and microornament. Dental plates are present briefly early in ontogeny. Based on the possession of transient dental plates, a septalium, and conspicuous microornament, Praeargyrotheca is removed from the family Megathyrididae and assigned, along with the two new genera, to a new family Praeargyrothecidae. Scanning electron microscopy study of loop development in both fossil and Recent terebratelloids and megathyrids supports a recent immunological study of living taxa from both groups that suggests that the two groups are not as closely linked phylogenetically as previously maintained. The family Megathyrididae is thus removed from the superfamily Terebratelloidea and assigned, along with the new family Praeargyrothecidae, to a new superfamily Megathyridoidea.


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