Genetic parameters for Australian maternal and dual-purpose meatsheep breeds. II. Liveweight, wool and reproduction in Corriedale sheep

1994 ◽  
Vol 45 (2) ◽  
pp. 469 ◽  
Author(s):  
LD Brash ◽  
NM Fogarty ◽  
AR Gilmour
Keyword(s):  
Litter Size ◽  
Genetic Correlations ◽  
Fibre Diameter ◽  
Dual Purpose ◽  
Scrotal Circumference ◽  
Average Fibre Diameter ◽  
Reproduction Traits ◽  
Component Traits ◽  
Average Fibre ◽  
Fleece Weight

Performance of sheep from a large Corriedale stud flock was assessed. Weaning and yearling liveweights, greasy and clean fleece weights, clean fleece yield and average fibre diameter records on approximately 3000 animals representing 64 sires were analysed. Also scrotal circumference of 1009 rams representing 27 sires were analysed. Some 3740 reproduction records over 11 years, for 980 ewes representing 114 sires, were also analysed. Heritability estimates were 0 34 �0.07 for weaning liveweight, 0.132 � 0.04 for yearling liveweight, 0.32 � 0.07 for greasy and 0.29 � 0.07 for clean fleece weights, 0.534 � 0 10 for clean fleece yield, 0-56 � 0 10 for yearling and 0 62 � 0 - 14 for hogget fibre diameters and 0.l5 � 0.11 for scrotal circumference. The genetic correlations between liveweight and fleece weight ranged from 0.13 to 0.34, and those for liveweight and fibre diameter were close to zero. The genetic correlations of scrotal circumference with liveweight and wool traits were positive and moderate. Reproductive performance in the flock was high, with fertility 92% and litter size 1.62, resulting in 148% lambs born and 130% lambs tagged alive of ewes joined. Estimated heritabilities for reproduction traits were 0-03 � 0.02 for lambs born and for lambs tagged, and the component traits, fertility 0 01 � 0 -02, litter size 0.04 � 0.03 and neonatal lamb survival was zero. Lambing date had a heritability of 0.06 � 0.03. Estimates of repeatability ranged from 0-04 � 0-01 for fertility to 0.07% 0.02 for lambs tagged and lambing date. Predicted heritabilities for the average of 3.8 records per ewe generally agreed with those estimated for average ewe lifetime performance, which ranged from 0.02 � 0.08 for fertility to 0.12 � 0.09 for lambs tagged and 0.l3 � 0.09 for lambing date.

Genetic parameters for reproduction and lamb production and their components and liveweight, fat depth and wool production in Hyfer sheep

1994 ◽  
Vol 45 (2) ◽  
pp. 443 ◽  
Author(s):  
NM Fogarty ◽  
LD Brash ◽  
AR Gilmour
Keyword(s):  
Genetic Correlation ◽  
Litter Size ◽  
Genetic Parameters ◽  
Genetic Correlations ◽  
Production Traits ◽  
Wool Production ◽  
Reproduction Traits ◽  
Component Traits ◽  
Lamb Survival ◽  
Fleece Weight

Hyfer is a new composite breed suited to specialized lamb production and developed from Dorset (+), Booroola Merino (+)and Trangie Fertility Merino (+) genotypes. Genetic parameters estimated by restricted maximum likelihood methods are reported for liveweight, wool production and a range of reproduction and lamb production traits. Estimates of heritability were 0 44 � 0.10, 0.222 � 0-08 and 0.28 � 0.08 for liveweight, ultrasonic fat depth (C site) and fat depth adjusted for liveweight respectively, for 1454 rams averaging 14 months of age, which were the progeny of 130 sires. The genetic correlation for ram liveweight and fat depth was 0.06 � 0.17 and -0.30 � 0.16 with adjusted fat depth. Estimates of heritability for ewe liveweight at 12-18 months of age and greasy fleece weight at 11-19 months of age were 0.61 � 0.10 and 0.38 � 0.08 respectively, with a genetic correlation of 0.35 � 0.13, based on 1619 ewes, which were the progeny of 135 sires. A maternal effect was significant for ewe liveweight and its inclusion in the model reduced the estimate of heritability to 0.44 � 0.09. Young ewes were joined three times in two years in an accelerated 8-monthly lambing system, and joinings commenced in mid-February, late October and mid-June. The estimates of heritability for overall ewe performance were for lambs born 0.09 � 0.03, lambs weaned 0.04 � 0.02 and weight of lamb weaned 0 -06 � 0.02 (all per ewe joined). Heritabilities for the component traits were: fertility 0 -03 � 0 02, litter size 0.19 � 0.04 and lamb survival 0.07 � 0.03. Estimates of repeatability for the corresponding traits ranged from 0.11 to 0.15, except for litter size (0.26 � 0.02). Estimates of heritability for lambs born, lambs weaned and weight weaned were higher for the first joining than when estimates were based on three joining records. This was associated with higher heritability for fertility from first joining (0.10 � 0.06) than for all three records and the reverse for lamb survival. Estimates of heritability for mean ewe performance were considerably higher and in agreement with the values predicted from the repeatability analyses and number of records, illustrating the value of using repeated ewe records for selection. The genetic correlations between the component traits and overall measures of lamb production were not high enough to consider using component traits as indirect selection criteria. The genetic correlations of liveweight with reproduction traits were positive, but generally not significant, and ranged from 0.01 to 0.63, except for litter size (-0.19 � 0.17). The genetic correlations of greasy fleece weight with reproduction traits were generally smaller than those for liveweight, were not significant and ranged from -0.12 to 0.32. These results, together with others, will be used to compile parameter sets for use in LAMBPLAN to provide meatsheep breeders with comprehensive breeding objectives incorporating growth, leanness, wool and reproduciton.

Genetic correlations between early growth and wool production of crossbred ewes and their subsequent reproduction

2009 ◽  
Vol 49 (1) ◽  
pp. 17 ◽  
Author(s):  
R. A. Afolayan ◽  
N. M. Fogarty ◽  
A. R. Gilmour ◽  
V. M. Ingham ◽  
G. M. Gaunt ◽  
...  
Keyword(s):  
Growth Traits ◽  
Genetic Correlations ◽  
Fibre Diameter ◽  
Reproductive Traits ◽  
Early Growth ◽  
Wool Production ◽  
Component Trait ◽  
Average Fibre Diameter ◽  
Average Fibre ◽  
Fleece Weight

Genetic correlations between reproductive and productivity traits of ewes and their early growth, wool production and worm resistance traits were estimated among 2460 crossbred ewes. The ewes were progeny of 74 maternal breed sires and mainly Merino dams. The ewes were born over 3 years and had three joinings with a total of 6824 ewe joining records. The reproductive and ewe productivity traits included: (i) fertility (ewes lambing per ewe joined); (ii) litter size (number of lambs born per ewe lambing); (iii) rearing ability or lamb survival (lambs weaned of lambs born for lambing ewes); (iv) number of lambs born (NLBj) per ewe joined; (v) number of lambs weaned (NLWj) per ewe joined; (vi) total litter weight weaned (TWWj) per ewe joined; and (vii) the component trait average lamb weaning weight in the litter (AWW). The growth traits included the weight of the ewe at birth (BWT), weaning (WWT) and postweaning (PWWT), as well as growth rate pre- and postweaning. The wool traits included greasy (GFW) and clean fleece weight (CFW), yield and average fibre diameter (FD) at their first adult shearing. Worm egg count (WEC) during their first year was also recorded. Bivariate mixed models analyses using ASReml procedures were used to estimate the genetic correlations. The genetic correlations between TWWj and growth traits were positive and moderate to high and ranged from 0.34 for BWT to 0.61 for PWWT. Those for the other composite reproductive traits were slightly lower and ranged from 0.27 to 0.54 for NLWj and from 0.07 to 0.46 for NLBj. The genetic correlations between the overall measures of reproduction (NLBj, NLWj and TWWj) and GFW, CFW and FD were negative and generally low to moderate (–0.21 to –0.52). The correlations were generally larger for NLBj than for NLWj and TWWj. The genetic correlations between the reproductive traits and WEC were close to zero, except between AWW and WEC (0.42). The phenotypic correlations between the reproductive traits and the growth, wool and WEC traits were all close to zero. The genetic correlations were generally favourable between ewe reproduction and growth, with early selection for traits such as WWT and PWWT likely to give some improvement in subsequent reproduction and ewe productivity. In contrast, there appears to be a slight antagonism between reproduction and wool production. This study provides estimates of genetic correlations from crossbred ewes that add to the limited knowledge of these parameters that will improve the accuracy of genetic evaluation and prediction of the outcomes from breeding programs for meat and wool objectives that include reproduction.

Genetic importance of fat and eye muscle depth in Merino breeding programs

2016 ◽  
Vol 56 (4) ◽  
pp. 690 ◽  
Author(s):  
D. J. Brown ◽  
A. A. Swan
Keyword(s):  
Carcass Traits ◽  
Genetic Correlations ◽  
Fibre Diameter ◽  
Economic Index ◽  
Parasite Resistance ◽  
Eye Muscle ◽  
Breeding Programs ◽  
Dual Purpose ◽  
Internal Parasite ◽  
Reproduction Traits

Australian Merino breeders have traditionally selected animals for breeding predominately on the basis of wool characteristics. Over recent decades, an increasing proportion of Merino breeders are interested in producing a ewe that can be used for prime-lamb production, but that also performs well for wool characteristics. Correlations between ultrasound carcass traits and other traits such as wool, internal parasite resistance and reproduction traits, are not very well known. The aims of this study were three-fold: (1) to estimate the genetic relationships between ultrasound carcass traits and wool, internal parasite resistance and reproduction traits, (2) to determine the value of recording ultrasound carcass traits in Merino breeding programs, and (3) to evaluate the impact of improving ewe genetic merit for fatness on their reproduction performance. Ultrasound fat and eye muscle depth had small to moderate genetic correlations with most traits, with positive correlations observed for bodyweight, fibre diameter, fibre curvature and reproduction, and negative correlations observed for fleece weight, fibre diameter coefficient of variation, worm egg count and breech wrinkle. As expected on the basis of these genetic correlations, estimated breeding values for fat depth of ewes had a positive association with their observed reproduction performance, but the effect explained only minimal variation in reproductive performance, and was extremely variable among flocks and years. A range of measurement scenarios was investigated for three standard MERINOSELECT indexes. Measuring fat and eye muscle depth resulted in 3%, 4% and 21% additional economic index gain for the fine, medium and dual purpose indexes, respectively, whereas measuring reproduction traits directly resulted in 17%, 27% and 45% additional gain in the economic index. Dual purpose index gains benefited more from measuring ultrasound carcass traits as it is the only index with a direct economic value placed on carcass traits. Measuring fat and eye muscle depth also resulted in a greater reduction in worm egg count. The results indicated that desirable genetic progress can be made in wool, ultrasound carcass, internal parasite resistance and number of lambs born and weaned simultaneously using multiple trait selection to account for the mix of favourable and unfavourable correlations between these traits. These results also demonstrated that the best method to maximise economic gain is to measure as many traits (or closely correlated traits) in the breeding objective as possible.

Studies on Western Australian Merino sheep. 1. Stud, strain and environmental effects on hogget performance

1992 ◽  
Vol 43 (6) ◽  
pp. 1381 ◽  
Author(s):  
RP Lewer ◽  
RR Woolaston ◽  
RR Howe
Keyword(s):  
Fibre Diameter ◽  
Strain Differences ◽  
Average Fibre Diameter ◽  
Weaning Age ◽  
Average Fibre ◽  
Males And Females ◽  
Wool Yield ◽  
Fleece Weight ◽  
Western Australian ◽  
Australian Merino

A 6 year study is reported of Merino studs and strains (Peppin, Collinsville and Bungaree) in Western Australia. Wool and body traits of males and females were measured, with additional subjective traits assessed on females. The effects of strain, stud (within strain), birth year, dam age, birth rearing rank, weaning age and their interactions were estimated using least squares procedures. When tested against studs, strain differences were significant for fibre diameter (both sexes), clean wool yield and about half of the subjective traits (females) but for none of the liveweights. Stud and year effects were significant for all traits, as was their interaction for most traits. Some studs were more stable between years than others in both clean fleece weight and average fibre diameter. Of the remaining effects, birth rearing rank influenced the greatest number of traits, while dam age only affected yield in ewes and some early liveweights. Peppins produced wool 2.0-2.3 microns finer than Bungarees, but not significantly different from Collinsvilles. Peppins also had the best subjective wool scores, but had the highest wrinkle scores and scored poorly on other subjective body traits. Twin-born hoggets produced 0.05-0.15 kg less clean wool than their single-born contemporaries, and their fleeces were about 0.4 microns coarser with poorer subjective qualities. Twins were also lighter from birth (by 23%) up to 17 months (by 5%) in females. Late-born lambs had higher birth weights, but lower subsequent weights, persisting until 12 months in females.

Sensitivity of breeding objectives to prices and genetic parameters in Australian Corriedale and Polwarth dual-purpose sheep

1993 ◽  
Vol 33 (3) ◽  
pp. 259 ◽  
Author(s):  
NM Fogarty ◽  
AR Gilmour
Keyword(s):  
Feed Intake ◽  
Selection Index ◽  
Genetic Correlations ◽  
Fibre Diameter ◽  
Dual Purpose ◽  
Wool Production ◽  
Price Differentials ◽  
Breeding Objective ◽  
Feed Costs ◽  
Fleece Weight

A profit equation was developed for Australian Corriedale and Polwarth dual-purpose sheep kept for both wool and lamb production. Ten traits contribute to income and costs and were included in the breeding objective. These encompass wool production, fibre diameter, and feed intake of breeding ewes and hogget ewe replacements; sale weight and carcass fat depth of lambs; and reproductive rate and mature weight of ewes. The relative economic value for each trait in the enterprise was calculated from the profit equation using income and costs for 1 year. Overall gain ($A) and gain in each trait in the breeding objective from selection using an index of hogget greasy fleece weight, hogget fibre diameter, dam's number of lambs weaned, lamb weight, and lamb fat depth are reported. The sensitivity of the genetic changes in each trait in the breeding objective and index coefficients were assessed for a range of prices of products and feed costs. Sensitivity to changes in heritabilities and genetic correlations was also assessed. The incorporation of these maternal traits into LAMBPLAN is discussed. For the standard parameters and prices used, gain in leanness accounted for one-third of overall gain ($/ewe. s.d. of selection). Other traits that contributed to overall gain were fibre diameter (28%), reproduction (18%), and growth (14%). There was a small gain through feed intake (8%), and a very small loss in wool weight. Measures of fatness, growth, and fibre diameter were the important traits in the selection index. Inclusion of hogget greasy fleece weight and dam's number of lambs weaned each added <1% to the efficiency of the selection index. Varying the sale price for lamb and the price differentials for fibre diameter and fat depth had the greatest impact on overall gain. High lamb price increased gains in both reproduction and lamb weight, whereas, high price differentials for fibre diameter and fat mainly increased gains in the particular trait. A large range in prices for wool had very little effect on the individual traits or overall. Changing feed costs had little effect on overall gain, although high feed cost reduced gains from reproduction which were compensated by reduced feed intake. Halving the heritability value for each trait reduced overall gain, largely through reduction in the trait. Varying the genetic correlations of wool production with other traits had little effect. However, when genetic correlations of reproduction with weight, fat, and fibre diameter were varied there were changes in overall gain, largely through reproduction.

The effects of breed, crossbreeding and other factors on variation in fleece and skin traits

1980 ◽  
Vol 30 (3) ◽  
pp. 417-429 ◽  
Author(s):  
J. A. Woolliams ◽  
G. Wiener
Keyword(s):  
Unit Area ◽  
Fibre Diameter ◽  
Fibre Length ◽  
Live Weight ◽  
Strong Relationship ◽  
Average Fibre Diameter ◽  
Wide Range ◽  
Average Fibre ◽  
Fleece Weight ◽  
The Relationship

ABSTRACTBreed and crossbreeding significantly affected variation in 10 fleece, follicle density and fibre diameter traits in 12-week-old lambs. The breeds involved were the Scottish Blackface, South Country Cheviot, Welsh Mountain, Lincoln Longwool, Southdown and Tasmanian Merino, but the last only by rams.The Southdown came closest to the Merino in affecting density and diameter traits although the Merino crosses remained clearly distinguishable by their high follicle density. The Lincoln and Blackface crosses were notable for their long fibres and the Blackface crosses also had the least dense follicle and coarsest primary fibres. The coarsest secondary fibres were from Lincoln and Cheviot crosses. Heterosis estimates were obtained for crosses involving the Blackface, Cheviot, Welsh and Lincoln breeds. Significant heterosis was present in a wide range of traits but was not apparent in all crosses. Maternal differences were evident for many of the traits but with density these differences were apparently related to maternal effects on live weight of the lamb. The sex of the lamb, its birth type and rearing and its 12-week live weight significantly affected many of the traits.Among breeds and crosses a strong relationship was found between fibre density and average fibre diameter. The relationship suggested that breed variation in total fibre area per unit area of skin was small compared with the breed variation in its components. Fleece weight per unit area was most closely related to mean fibre length.

Wool yield and characteristics of Dorset, Leicester and Sunffolk breeds and their ‘DLS’ cross

1977 ◽  
Vol 88 (3) ◽  
pp. 651-653 ◽  
Author(s):  
M. H. Fahmy ◽  
J. A. Vasely
Keyword(s):  
First Generation ◽  
Fibre Diameter ◽  
Fibre Length ◽  
Wool Production ◽  
Average Fibre Diameter ◽  
Average Fibre ◽  
Wool Yield ◽  
Fleece Weight

SUMMARYGreasy fleece weight (12 months growth) and wool samples from hip and shoulder regions were taken on 15 Dorset, 20 Leicester, 20 Suffolk and 17 DLS ewes (a cross of ½ Dorset, ¼ Leicester, ¼ Suffolk obtained by mating DL rams to DS ewes and DS rams to DL ewes). The purpose of the study was to compare wool production and characteristics of the first generation of DLS with that of the three breeds of origin. Wool production of DLS was 3·58 kg, 8·8 and 14% higher than that of Suffolk and Dorset, but 11% lower than that of Leicester (P< 0·01). The percentage of clean wool was highest in Leicester (78·3%), followed by DLS, Dorset and Suffolk (76·1, 74·3 and 70·5%, respectively). The average fibre diameter of the DLS and Leicester was 38 /m, 4 /m thicker than that for Suffolk and Dorset (P < 0·01). Average fibre length was 15 cm in DLS and Dorset, 6 cm shorter than in Leicester, and 3 cm longer than in Suffolk (P < 0·01). The variability in fibre diameter and length was highest in Leicester, followed closely by DLS.

Genetic studies on growth, reproduction and wool productiontraits in Harnali sheep

2016 ◽  
Author(s):  
SPACE Lalit ◽  
Z. S. Malik ◽  
D. S. Dalal ◽  
C. S. Patil ◽  
S. P. Dahiya
Keyword(s):  
Body Weight ◽  
Growth Traits ◽  
Additive Genetic Variance ◽  
Genetic Correlations ◽  
Fibre Diameter ◽  
Reproductive Traits ◽  
Phenotypic Correlation ◽  
Reproduction Traits ◽  
Heritability Estimates ◽  
Fleece Weight

Data on growth, reproduction and wool traits of 1603 Harnali sheep maintained at Lala Lajpat Rai University of Veterinary and Animal Sciences, Hisar over a period of 22 years (1992-2013) were utilized for genetic analysis. The overall least squares mean for birth weight (BW), weaning weight (WW), six month body weight (SMW), age at first lambing (AFL), weight at lambing (WL), average lambing interval (ALI), greasy fleece weight (GFW), staple length (SL) and Fibre diameter (FD) were estimated as 3.35±0.02 kg, 12.41±0.08 kg, 16.30±0.12 kg, 707.05±2.07 days, 26.91±0.10 kg and 402.85±2.40 days, 1.62±0.02 kg, 5.65±0.03 cm and 25.85±0.07 μ, respectively. The effect of year of birth, sex of lamb and dam's weight at lambing were significant for all growth traits. The effect of year of birth and dam's weight at lambing were significant for all reproduction traits and GFW. No definite trend was observed over the years for body weights and reproductive traits. The effect of sex was significant for early growth traits. The heritability estimates were moderate for all the growth traits with high genetic correlations of BW and WW with SMW. Reproduction traits had lower estimates of heritability which indicated presence of lower additive genetic variance for these traits. Heritability estimates for studied wool traits were moderate to high. Positive genetic and phenotypic correlation of BW and WW with six month body weight and grease fleece weight indicated that selection for six month body weight would increase body weight and grease fleece weight.

Reproductive performance and genetic parameters for Australian Dorset Sheep

1994 ◽  
Vol 45 (2) ◽  
pp. 427 ◽  
Author(s):  
LD Brash ◽  
NM Fogarty ◽  
AR Gilmour
Keyword(s):  
Litter Size ◽  
Reproductive Performance ◽  
Genetic Improvement ◽  
Genetic Correlations ◽  
Reproductive Traits ◽  
Reproductive Rate ◽  
Parameter Estimates ◽  
Reproduction Traits ◽  
Component Traits ◽  
Lamb Survival

Reproductive performance of ewes in a large Dorset stud flock is reported. The data included 2488 ewes with 10016 joining records over 25 years. Average performance for reproductive traits were: fertility 92%, litter size 1.31, neonatal lamb survival 91%, lamb survival to weaning 85%, with overall lambs born 1.20 and lambs weaned 1.02 per ewe joined. Differences between breeds (Poll Dorset and Dorset Horn), season of birth (spring and autumn) and season of joining (spring and autumn) were significant for most reproduction traits. Linear and quadratic regressions for age of ewe at joining were highly significant for all traits (P < 0.001) with maximum performance between three and six years of age. Estimates of heritability for ewe reproductive traits were: 0.062 � 0-02 for number of lambs born and 0.04 � 0.01 for number of lambs weaned per ewe joined, and component traits, 0.02 �0 -01 for fertility, 0 08 � 0 -02 for litter size and 0.00 � 0.01 for lamb survival. Estimates of repeatability were less than 0.14 for all the reproduction traits, which resulted in substantially higher predicted heritabilities when repeated records were used. Heritability estimates for average ewe lifetime performance, with approximately four records, were 0.08 � 0 06 for lambs born, 0.12 � 0.05 for lambs weaned, 0.08 � 0.04 for fertility, 0.l9 � 0 -04 for litter size and zero for lamb survival. The genetic correlations between litter size and lambs born and weaned were close to unity, whereas those for fertility were lower and declined from lambs born (0.6l � 0.22) to lambs weaned (0-45 � 0.30). The estimated heritability for ewe longevity, defined as the number of years the ewe remained in the breeding flock, was 0.00 � 0.03. The potential for genetic improvement in reproductive rate is discussed in relation to other reports in the literature and in the context of the roles of the Dorset breed in the Australian lamb industry. Increased reproductive rate in Dorset flocks impacts on stud profitability and the rate of genetic improvement from selection for other traits such as liveweight. The parameter estimates derived will be used in compiling breed-specific parameter sets for genetic evaluation in LAMBPLAN.

Productivity of classers' grades and sire selection differentials for fleece weight in two Merino studs

1982 ◽  
Vol 22 (117) ◽  
pp. 274
Author(s):  
BJ McGuirk ◽  
M Rose ◽  
R Scott
Keyword(s):  
Fibre Diameter ◽  
Selection Procedure ◽  
The Other ◽  
Average Fibre Diameter ◽  
Average Coefficient ◽  
Average Fibre ◽  
Selection For ◽  
Fleece Weight ◽  
Selection Differentials ◽  
Sire Selection

The normal practice in Merino studs is to class the rams visually into three grades: reserve rams from which the stud sires will be chosen; flock rams for use in commercial flocks; and culls. This study examined the productivity of these grades in three drops of rams from each of two studs. In each drop there were differences between the grades in greasy and clean fleece weight. In stud 1 the relative clean and greasy wool percentages in the reserve, flock and cull grades were 11 6,98 and 86, and 11 6,98 and 87 (P < 0.05), respectively. Clean fleece weights were not available on the visual culls in stud 2. In that stud the relative greasy wool percentages of rams in three grades were 112, 98 and 90, and the reserve rams also cut 15% more clean wool than the flock rams (P< 0.05). The differences between the grades in yield and average fibre diameter were less than they were for fleece weight and generally not statistically significant. In stud 1 the reserve rams were 0.6% higher yielding and 0.2 pm stronger than the drop average. The reserve rams were less variable than the other grades in fleece weights, yield and diameter. The largest differences were for fleece weights, where the differences in variation between the grades were significant. The average coefficient of variation for greasy fleece weight among the reserve rams in the two studs was 10.0% compared with the estimate of 13.2% for the whole drop. Selection differentials for fleece weight were estimated for stud sires selected from these six drops of rams. The efficiency of sire selection for either greasy or clean fleece weight was in excess of 70% of the maximum possible. In the four drops in which all sire replacements were selected exclusively from the reserves, the two steps in the sire selection procedure contribution almost equally to the sire selection differentials achieved.

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