The Occurrence of Matairesinol in Mountain Hemlock (Tsuga mertensiana), Western Hemlock (Tsuga heterophylla), and Balsam (Abies amabilis)1

1962 ◽  
Vol 27 (1) ◽  
pp. 322-323 ◽  
Author(s):  
G. M. Barton ◽  
J. A. F. Gardner
Keyword(s):  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Abies Amabilis ◽  
Tsuga Mertensiana

STUDIES IN FOREST PATHOLOGY: VII. DECAY IN WESTERN HEMLOCK AND FIR IN THE FRANKLIN RIVER AREA, BRITISH COLUMBIA

1949 ◽  
Vol 27c (6) ◽  
pp. 312-331 ◽  
Author(s):  
D. C. Buckland ◽  
R. E. Foster ◽  
V. J. Nordin
Keyword(s):  
British Columbia ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Forest Pathology ◽  
Abies Amabilis ◽  
Armillaria Mellea ◽  
Forest Region ◽  
Volume Increment ◽  
Juan De Fuca

An investigation of decay in western hemlock (Tsuga heterophylla (Raf.) Sarg.) and fir (mainly Abies amabilis (Loud.) Forb.) in the Juan de Fuca forest region of British Columbia has shown that the major organisms causing root and butt rots are the same in both species. These are Poria subacida (Peck) Sacc., Fomes annosus (Fr.) Cke., Armillaria mellea Vahl ex Fr., Polyporus sulphureus Bull. ex Fr., and P. circinatus Fr. Those organisms causing trunk rots of western hemlock, in decreasing order of importance, are Fomes pinicola (Sw.) Cke., F. Pini (Thore) Lloyd, Stereum abietinum Pers., Fomes Hartigii (Allesch.) Sacc. and Trav., and Hydnum sp. (H. abietis). These same organisms causing trunk rots of fir, in decreasing order of importance, are Fomes pinicola, Stereum abietinum, Hydnum sp. (H. abietis), Fomes Pini, and Fomes Hartigii. The logs of 963 western hemlock were analyzed in detail. Maximum periodic volume increment was reached between 225 and 275 years of age. Maximum periodic volume increment was reached between 275 and 325 years of age in the 719 fir that were analyzed. Scars were the most frequent avenue of entrance for infection. In 59% of the cases of infection studied the fungus had entered through wounds.

REVISION OF SOME DIMERIELLA AND DIMEROSPORIUM PARASITES OF CONIFERS

1965 ◽  
Vol 43 (6) ◽  
pp. 631-639 ◽  
Author(s):  
R. A. Shoemaker
Keyword(s):  
North America ◽  
Abies Alba ◽  
Tsuga Canadensis ◽  
Tsuga Heterophylla ◽  
Eastern North America ◽  
Western North America ◽  
Abies Grandis ◽  
Abies Amabilis ◽  
Conifer Needles ◽  
Tsuga Mertensiana

Study of the type of Dimerosporium balsamicola (Peck) Ell. & Ev. (≡ Meliola balsamicola Peck) revealed that this species is a melioline hyperparasite. The fungus is transferred to Dimerium, redescribed, illustrated, and distinguished from Phacocryptopus nudus (Peck) Petrak, Asterina sp., and from Epipolaeum abietis (Dearness) n. comb. (≡ Dimerosporium abietis Dearness), which also occur on Abies in North America. In all, four species of Epipolaeum that occur on conifer needles are treated: Epipolaeum abietis on Abies grandis (Dougl.) Lindl. and Abies amabilis (Dougl.) Forb. in western North America, on Abies alba Mill. in France, and on Keteleeria davidiana Beiss. in Formosa; Epipolaeum tsugae (Dearness) n. comb. on Tsuga canadensis (L.) Carr. in eastern North America, and on Tsuga heterophylla (Raf.) Sarg. and Tsuga mertensiana (Bong.) Carr. in western North America; Epipolaeum pseudotsugae (V. M. Miller & Bonar) n. comb. on Pseudotsuga taxifolia (Poir.) Britton from western North America; Epipolaeum terrieri (Petrak in Terrier) n. comb. on Abies alba in Switzerland.

Pollen morphology and development of the pollination mechanism in Tsuga heterophylla and T. mertensiana

1983 ◽  
Vol 61 (12) ◽  
pp. 3041-3048 ◽  
Author(s):  
John N. Owens ◽  
Margaret Dianne Blake
Keyword(s):  
Reproductive Biology ◽  
Pollen Morphology ◽  
Pollen Tubes ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Pollination Mechanism ◽  
Pollination Drop ◽  
Tsuga Mertensiana

Pollen of Tsuga heterophylla (Raf.) Sarg. (western hemlock) is nonsaccate and bears spines, whereas pollen of Tsuga mertensiana (Bong.) Carr. (mountain hemlock) is saccate and lacks spines. The pollination mechanism in western hemlock consists of a short funnel-like integument tip with a large micropyle. Pollen may enter the micropyle or germinate on the bract or ovuliferous scale and form long pollen tubes. The pollination mechanism in mountain hemlock consists of two large micropylar flaps which secrete minute droplets to which pollen adheres. A pollination drop is not formed in either species. These features are discussed in relation to the taxonomy of the genus and the reproductive biology of these two species.

Above-ground biomass predicts growth limitation in amabilis fir and western hemlock seedlings

2003 ◽  
Vol 79 (2) ◽  
pp. 285-290 ◽  
Author(s):  
Alan K Mitchell ◽  
B Glen Dunsworth ◽  
Tom Bown ◽  
Jonathan A Moran
Keyword(s):  
Nitrogen Concentration ◽  
Volume Growth ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Stem Volume ◽  
Above Ground Biomass ◽  
Growth Limitation ◽  
Foliar Nitrogen ◽  
Abies Amabilis ◽  
Ground Biomass

Conifer regeneration on clearcut montane sites is frequently affected by post-planting growth stagnation. The ability to predict such stagnation would be a valuable asset to forest managers. In this study, we tested the usefulness of above-ground biomass and photosynthetic efficiency (as estimated by foliar nitrogen concentration) in diagnosing growth limitations in western hemlock (Tsuga heterophylla) and amabilis fir (Abies amabilis). Seedlings were grown under different silvicultural systems (clearcut, patch cut, green tree retention and shelterwood) and post-planting treatments (fertilization, vegetation removal) at the Montane Alternative Silvicultural Systems (MASS) site on Vancouver Island, BC. Foliar nitrogen was found to be a poor predictor of height and stem volume growth. However, above-ground biomass predicted current height and stem volume (year 3 after planting), as well as future stem volume (year 7 after planting), in both species. Above-ground biomass therefore represents a useful measure of likely future growth performance, and may provide early warning of incipient growth stagnation in these species. Key words: amabilis fir, Abies amabilis, biomass, western hemlock, Tsuga heterophylla, growth limitation, photosynthetic capacity, biomass, seedling growth, regeneration

Growth limitations of planted conifers regenerating under Montane Alternative Silviculture Systems (MASS): Seven-year results

2004 ◽  
Vol 80 (2) ◽  
pp. 241-250 ◽  
Author(s):  
A K Mitchell ◽  
B G Dunsworth ◽  
J T Arnott ◽  
R. Koppenaal ◽  
R. Benton ◽  
...  
Keyword(s):  
Treatment Time ◽  
Tsuga Heterophylla ◽  
Height Growth ◽  
Western Hemlock ◽  
Initial Growth ◽  
Vegetation Control ◽  
Abies Amabilis ◽  
Silvicultural System ◽  
The Impact ◽  
Effect On Growth

As part of the Montane Alternative Silviculture Systems (MASS) project, this study investigates limits on the growth of montane conifers resulting from varying overstory retention under conventional and alternative silvicultural systems. After harvesting treatments were complete in 1993, Abies amabilis (amabilis fir) and Tsuga heterophylla (western hemlock) seedlings were spring planted in replicated blocks of shelterwood (SW), patch cut (PC), green tree retention (GT) and clearcut (CC) systems. In addition, sub-plots were established within each silvicultural system in which fertilization (at planting) and vegetation control post-planting treatments were applied alone and in combination to test the extent to which growth limitations are related to nutrient availability and vegetative competition. The impact of overstory retention was most pronounced in the reduced light environment of the SW where height growth after seven years was 26-30% lower in both species compared to the untreated CC, GT and PC systems. Although the effect on growth of both species in the SW was mitigated somewhat by fertilization and vegetation control treatments, amabilis fir did not attain free-to-grow height (1.3 m) regardless of post-planting treatment. Time to free-to-grow height in the more open silvicultural systems was reduced in both fir and hemlock with fertilization and vegetation control alone and in combination, except in the CC where the initial growth response to fertilization alone was diminished by the end of the seven-year study. In contrast to fertilization, the effects of vegetation control on height growth were not apparent until three to five years and seven years after planting in the CC and GT, PC, SW, respectively. Combining vegetation control and fertilization had an additive effect on growth in amabilis fir but not in western hemlock. The effect of silvicultural systems and post-planting treatments on the two species illustrate that both above- and below-ground resource availability (light and nutrients) availability was potentially limiting to growth, particularly in the shelterwood treatment. Key words: MASS, silviculture systems, regeneration, Tsuga heterophylla, Abies amabilis, shelterwood, patch cut, green tree reten-tion, clearcut, fertilizer, vegetation control

Studies on periderm. II. Anthocyanidins in secondary periderm tissue of amabilis fir, grand fir, western hemlock, and western red cedar

1969 ◽  
Vol 47 (9) ◽  
pp. 1419-1422 ◽  
Author(s):  
D. B. Mullick
Keyword(s):  
Tsuga Heterophylla ◽  
Free State ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Abies Grandis ◽  
Abies Amabilis ◽  
Red Cedar ◽  
Western Red Cedar ◽  
Thuja Plicata Donn

By using a mild method, which precludes the possibility of pigment hydrolysis during extraction and processing, the occurrence of anthocyanidins in the free state has been detected in the normal secondary periderm tissues (rhytidomal region) of amabilis fir (Abies amabilis (Dougl.) Forbes), grand fir (Abies grandis (Dougl.) Lindl.), western hemlock (Tsuga heterophylla (Raf.) Sarg.), and western red cedar (Thuja plicata Donn.). Cyanidin was present in all species. Pelargonidin was present in western hemlock and most likely also in western red cedar.

Growth ofPseudotsuga menziesiiandTsuga heterophyllaseedlings along a light gradient: resource allocation and morphological acclimation

1997 ◽  
Vol 75 (9) ◽  
pp. 1424-1435 ◽  
Author(s):  
D. Mailly ◽  
J. P. Kimmins
Keyword(s):  
Leaf Morphology ◽  
Biomass Accumulation ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Low Light ◽  
Growing Seasons ◽  
Light Levels ◽  
Light Gradient ◽  
Relative Light Intensity

Silvicultural alternatives that differ in the degree of overstory removal may create shady environments that will be problematic for the regeneration of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco). Gradients of light in the field were used to compare mortality, growth, and leaf morphological acclimation of two conifer species of contrasting shade tolerances: Douglas-fir and western hemlock (Tsuga heterophylla (Raf.) Sarg.). Results after two growing seasons indicated that Douglas-fir mortality occurred mainly at relative light intensity (RLI) below 20%, while western hemlock mortality was evenly distributed along the light gradient. Height, diameter, and biomass of the planted seedlings increased with increasing light for both species but at different rates, and maximum biomass accumulation always occurred in the open. Douglas-fir allocated more resources to stem biomass than western hemlock, which accumulated more foliage biomass. Increases in specific leaf area for Douglas-fir seedlings occurred at RLI ≤ 0.4 and red/far red (R/FR) ratio ≤ 0.6, which appear to be the minimal optimum light levels for growth. Conversely, western hemlock seedlings adjusted their leaf morphology in a more regular pattern, and changes were less pronounced at low light levels. These results, along with early mortality results for Douglas-fir, suggest that the most successful way to artificially regenerate this species may be by allowing at least 20% of RLI for ensuring survival and at least 40% RLI for optimum growth. Key words: light, light quality, leaf morphology, acclimation.

TOXICITY OF SMOKE FROM SPOROPHORES OF THE INDIAN PAINT FUNGUS AND WOOD FROM WESTERN HEMLOCK TO YELLOWFEVER MOSQUITOES

1987 ◽  
Vol 119 (12) ◽  
pp. 1109-1115
Author(s):  
W.P.L. Osborn ◽  
J.H. Borden
Keyword(s):  
British Columbia ◽  
Aedes Aegypti ◽  
Combustion Products ◽  
Tsuga Heterophylla ◽  
Water Soluble ◽  
Western Hemlock ◽  
Aqueous Extracts ◽  
Smoke Water

AbstractTo mitigate the effects of mosquitoes, settlers in the Revelstoke area of British Columbia reportedly burned the sporophores of the Indian paint fungus, Echinodontium tinctorium (Ell. & Ev.) Ell. & Ev., a pathogen of western hemlock, Tsuga heterophylla (Raf.) Sarg., and true firs, Abies spp. Larval and adult yellowfever mosquitoes, Aedes aegypti (L.), were exposed to aqueous extracts of smoke (smoke-waters) from E. tinctorium sporophores, and from western hemlock sapwood and heartwood. Smoke-waters were of approximately equal toxicity to larvae. Fungus smoke-water, but not sapwood or heartwood smoke-waters, lost 50% of its potency in 5 months. Vapors from fungus smoke-water were significantly more toxic to adult mosquitoes than those from sapwood or heartwood. Thus smoke from E. tinctorium sporophores and T. heterophylla wood apparently contain different water-soluble combustion products toxic to A. aegypti.

Genotype × shade effects for western hemlock

2005 ◽  
Vol 35 (6) ◽  
pp. 1496-1501
Author(s):  
G R Johnson ◽  
C Cartwright
Keyword(s):  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Different Levels

Western hemlock (Tsuga heterophylla (Raf.) Sarg.) families were grown under different levels of shade for 2 or 3 years at two nursery sites to determine whether families performed differently relative to one another in the different shade environments. Differences were found both for levels of shade and families, but no family × shade interaction was found. Results suggest that families selected in full-sun environments (clearcuts or farm fields) may be well suited for use in silvicultural systems where seedlings are planted in understory conditions.

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