scholarly journals A Problematic Tyrannosaurid (Dinosauria: Theropoda) Skeleton and Its Implications for Tyrannosaurid Diversity in the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta

2019 ◽  
Vol 303 (4) ◽  
pp. 673-690 ◽  
Author(s):  
Jordan C. Mallon ◽  
Jonathan R. Bura ◽  
Dirk Schumann ◽  
Philip J. Currie
Keyword(s):  
Upper Cretaceous ◽  
Horseshoe Canyon Formation

Quantitative diagenetic analyses of Edmontosaurus regalis (Dinosauria: Hadrosauridae) postcranial elements from the Danek Bonebed, Upper Cretaceous Horseshoe Canyon Formation, Edmonton, Alberta, Canada: implications for allometric studies of fossil organisms

2014 ◽  
Vol 51 (11) ◽  
pp. 1007-1016 ◽  
Author(s):  
Mackenzie Baert ◽  
Michael E. Burns ◽  
Philip J. Currie
Keyword(s):  
Cross Section ◽  
Upper Cretaceous ◽  
Strong Relationship ◽  
Major Axis ◽  
Minor Axis ◽  
Morphological Features ◽  
Fossil Assemblages ◽  
Fossil Vertebrate ◽  
Horseshoe Canyon Formation ◽  
Upper Campanian

For fossil assemblages, quantitative size and shape studies are often complicated by diagenetic distortion. Different vertebrate elements, although subjected to similar burial stresses, exhibit deformations based upon their original shapes; this hypothesis is tested here by quantitatively comparing deformed humeri and femora from the Danek Bonebed (a monodominant Edmontosaurus regalis bonebed from the upper Campanian Horseshoe Canyon Formation in Edmonton, Alberta, Canada) with samples of undeformed humeri and femora from modern and fossil assemblages. Analyses suggest that at the Danek Bonebed a strong relationship exists between element length and circumference despite being distorted by crushing deformation. Major and minor axes of the midshaft cross section, however, were not uniformly distorted. Although their anatomical position did not change, the major axis became longer relative to the minor axis in distorted specimens. A regression based on the undeformed humeri was not able to accurately predict circumference in the Danek humeri. Further study might quantify the deformation of other bones in the Danek Bonebed and could be extended to other assemblages and genera. Caution should be taken when conducting studies in which diagenetic crushing may have altered morphological features of fossil vertebrate remains.

Histologic growth dynamic study of Edmontosaurus regalis (Dinosauria: Hadrosauridae) from a bonebed assemblage of the Upper Cretaceous Horseshoe Canyon Formation, Edmonton, Alberta, Canada

2014 ◽  
Vol 51 (11) ◽  
pp. 1023-1033 ◽  
Author(s):  
Evan Vanderven ◽  
Michael E. Burns ◽  
Philip J. Currie
Keyword(s):  
Life History ◽  
Body Size ◽  
Upper Cretaceous ◽  
Growth Dynamics ◽  
Growth Dynamic ◽  
History Data ◽  
Life History Data ◽  
Fundamental Systems ◽  
Horseshoe Canyon Formation ◽  
Upper Campanian

The Danek Bonebed (Edmonton, Alberta, Canada) is a monodominant Edmontosaurus regalis assemblage of the upper Campanian (Upper Cretaceous) Horseshoe Canyon Formation. Bone histology of humeri and femora are used in this paper to test hypotheses about the growth dynamics and palaeobiology of Edmontosaurus. The high number of elements collected from the Danek Bonebed allow for an expansion of the multi-element histological record for hadrosaurs. Results indicate that Edmontosaurus had a growth trajectory similar to other large-bodied dinosaurs and reached the onset of somatic maturity at about 10–15 years of age; however, even the largest elements to preserve lines of arrested growth do not have external fundamental systems. This timing of the onset of somatic maturity agrees with the estimated body size of Edmontosaurus relative to other dinosaurs for which life-history data are available. Vascularity patterns support the hypothesis that edmontosaurs preserved at the Danek Bonebed were not subject to the same extreme seasonal environmental shifts as congenerics preserved at higher latitudes, further supporting overwintering behaviour in the latter.

High-precision U–Pb CA–ID–TIMS dating and chronostratigraphy of the dinosaur-rich Horseshoe Canyon Formation (Upper Cretaceous, Campanian–Maastrichtian), Red Deer River valley, Alberta, Canada

2020 ◽  
Vol 57 (10) ◽  
pp. 1220-1237 ◽  
Author(s):  
David A. Eberth ◽  
Sandra L. Kamo
Keyword(s):  
High Precision ◽  
Coastal Plain ◽  
Upper Cretaceous ◽  
Red Deer ◽  
Alluvial Plain ◽  
Type Area ◽  
Southern Alberta ◽  
Dinosaur Evolution ◽  
Horseshoe Canyon Formation ◽  
Upper Campanian

The non-marine Horseshoe Canyon Formation (HCFm, southern Alberta) yields taxonomically diverse, late Campanian to middle Maastrichtian dinosaur assemblages that play a central role in documenting dinosaur evolution, paleoecology, and paleobiogeography leading up to the end-Cretaceous extinction. Here, we present high-precision U–Pb CA–ID–TIMS ages and the first calibrated chronostratigraphy for the HCFm using zircon grains from (1) four HCFm bentonites distributed through 129 m of section, (2) one bentonite from the underlying Bearpaw Formation, and (3) a bentonite from the overlying Battle Formation that we dated previously. In its type area, the HCFm ranges in age from 73.1–68.0 Ma. Significant paleoenvironmental and climatic changes are recorded in the formation, including (1) a transition from a warm-and-wet deltaic setting to a cooler, seasonally wet-dry coastal plain at 71.5 Ma, (2) maximum transgression of the Drumheller Marine Tongue at 70.896 ± 0.048 Ma, and (3) transition to a warm-wet alluvial plain at 69.6 Ma. The HCFm’s three mega-herbivore dinosaur assemblage zones track these changes and are calibrated as follows: Edmontosaurus regalis – Pachyrhinosaurus canadensis zone, 73.1–71.5 Ma; Hypacrosaurus altispinus – Saurolophus osborni zone, 71.5–69.6 Ma; and Eotriceratops xerinsularis zone, 69.6–68.2 Ma. The Albertosaurus Bonebed — a monodominant assemblage of tyrannosaurids in the Tolman Member — is assessed an age of 70.1 Ma. The unusual triceratopsin, Eotriceratops xerinsularis, from the Carbon Member, is assessed an age of 68.8 Ma. This chronostratigraphy is useful for refining correlations with dinosaur-bearing upper Campanian–middle Maastrichtian units in Alberta and elsewhere in North America.

scholarly journals The first record of dinosaur eggshell from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada

2018 ◽  
Author(s):  
Gregory Funston ◽  
Philip J. Currie
Keyword(s):  
Upper Cretaceous ◽  
First Record ◽  
External Layer ◽  
A Site ◽  
Horseshoe Canyon Formation

Eggs and eggshell are generally rare in the Upper Cretaceous rocks of Alberta, despite being relatively abundant nearby in Montana. Palaeontologists and other people have been prospecting the Horseshoe Canyon Formation for more than a 130 years, but eggshell fragments have only just been recovered. The fragments are unornamented with angusticanaliculate pores and three structural layers. Numerous features support their referral to Prismatoolithus levis, and they confirm the presence of a bird-like external layer in this ootaxon. The fragments, which likely belonged to Albertavenator curriei, are from a site with abundant troodontid teeth and perinate material from hadrosaurs, ceratopsians, and theropods. The discovery of eggshell challenges the notion that the Horseshoe Canyon Formation is too heavily sideritized to preserve eggshell.

New information on the rare horned dinosaur Arrhinoceratops brachyops (Ornithischia: Ceratopsidae) from the Upper Cretaceous of Alberta, Canada

2014 ◽  
Vol 51 (6) ◽  
pp. 618-634 ◽  
Author(s):  
Jordan C. Mallon ◽  
Robert Holmes ◽  
Jason S. Anderson ◽  
Andrew A. Farke ◽  
David C. Evans
Keyword(s):  
Phylogenetic Analysis ◽  
Missing Data ◽  
Upper Cretaceous ◽  
Horizontal Section ◽  
New Information ◽  
Lateral Ridge ◽  
Minimum Age ◽  
Age Range ◽  
Horseshoe Canyon Formation

Arrhinoceratops brachyops is a poorly understood chasmosaurine ceratopsid from the Upper Cretaceous Horseshoe Canyon Formation of Alberta, previously described on the basis of only a single skull. Here, we report on a second specimen attributable to this species, including a relatively complete skull, syncervical, and partial left forelimb. This second specimen clarifies aspects of morphology not visible in the holotype, and also elucidates variation in A. brachyops. The species is distinguished by a square-shaped triangular process of the premaxilla, a steeply inclined triturating surface of the predentary, and a triangular nasal horncore in horizontal section. The dentary is also distinctive in bearing a bony lateral ridge similar to that of Anchiceratops ornatus, but more strongly developed. Phylogenetic analysis cannot resolve the relationships of Arrhinoceratops beyond the level of Chasmosaurinae, owing to both missing data and conflicting characters. However, we do find some support for a deep split within Chasmosaurinae, contrary to conventional topologies. We also report on other fragmentary specimens plausibly attributable to A. brachyops that suggest a minimum age range of approximately 750 ka for this species.

Palaeopathological changes in a population of Albertosaurus sarcophagus from the Upper Cretaceous Horseshoe Canyon Formation of Alberta, CanadaThis article is one of a series of papers published in this Special Issue on the theme Albertosaurus.

2010 ◽  
Vol 47 (9) ◽  
pp. 1263-1268 ◽  
Author(s):  
Phil R. Bell
Keyword(s):  
Upper Cretaceous ◽  
Unknown Origin ◽  
Limited Range ◽  
Special Issue ◽  
Puncture Wound ◽  
The Past ◽  
Bite Marks ◽  
Minimum Number ◽  
Lower Maastrichtian ◽  
Horseshoe Canyon Formation

Over the past decade, excavations of a monodominant theropod bonebed from the lower Maastrichtian beds of the Horseshoe Canyon Formation have recovered a minimum number of 26 individuals of the tyrannosaur, Albertosaurus sarcophagus . Examination of skeletal elements from the bonebed revealed a small number of abnormalities from at least two individuals. These include bony spurs (enthesophytes) of unknown origin on three pedal phalanges. Well-healed fracture calluses are present on two rib shafts and a gastralia element. The left dentary of one adult preserves both healed and unhealed parallel bite marks and a perforative lesion attributable to a partially healed, mechanically induced puncture wound. Unfortunately, the limited range in the types and frequencies of pathological changes provides only little information on the distribution of such phenomena but may be suggestive of the overall “health” of the population.

Enhancing bonebed mapping with GIS technology using the Danek Bonebed (Upper Cretaceous Horseshoe Canyon Formation, Edmonton, Alberta, Canada) as a case study

2014 ◽  
Vol 51 (11) ◽  
pp. 987-991 ◽  
Author(s):  
Katherine Bramble ◽  
Michael E. Burns ◽  
Philip J. Currie
Keyword(s):  
Information System ◽  
Upper Cretaceous ◽  
Gis Technology ◽  
Digital Map ◽  
Anatomical Information ◽  
The City ◽  
Horseshoe Canyon Formation ◽  
Specimen Identification ◽  
Upper Campanian

The Danek Bonebed is a monodominant Edmontosaurus bonebed preserving predominantly disarticulated material from the Upper Campanian Horseshoe Canyon Formation within the city limits of Edmonton, Alberta, Canada. To date, at least six dinosaur taxa have been identified at the site: Albertosaurus sarcophagus, Chasmosaurinae indet., Dromaeosauridae indet., Edmontosaurus regalis, Ornithomimidae indet., and Troodontidae indet. This bonebed has been used as a case study for creating a digital, searchable bonebed map using a geographic information system (GIS) platform. The original quarry maps produced on site are refined when digitized with new anatomical information gathered during preparation of collected specimens. Each specimen is labeled with the known specimen identification, quarry coordinates, and catalogue number. Creating a digital map of the bonebed allows easier interpretation of data and the ability to share maps to compare specific elements within the bonebed.

Upper Cretaceous palynostratigraphy of the Dry Island areaThis article is one of a series of papers published in this Special Issue on the theme Albertosaurus.

2010 ◽  
Vol 47 (9) ◽  
pp. 1145-1158 ◽  
Author(s):  
E. B. Koppelhus ◽  
D. R. Braman
Keyword(s):  
Large Scale ◽  
Upper Cretaceous ◽  
Colour Change ◽  
Special Issue ◽  
Significant Events ◽  
Plant Groups ◽  
Pollen Species ◽  
Lower Maastrichtian ◽  
Horseshoe Canyon Formation ◽  
Important Marker

The overall palynology of the upper Horseshoe Canyon Formation to lower Scollard Formation within the Edmonton Group is reviewed providing new insights into biostratigraphic relationships for the interval, including the recognition of unconformities in the stratigraphic package. The palynomorph recovery from the Horseshoe Canyon Formation and overlying units documents a number of significant events that allow the subdivision of the section. The Albertosaurus bonebed occurs within the lower Maastrichtian portion of the Edmonton Group. The bonebed is situated just below the base of the Scollardia trapaformis Palynomorph Zone, an important marker assemblage that has been documented from areas of the Yukon, Northwest Territories, Alberta, and Saskatchewan. This indicates that the bonebed lies at the top of the Mancicorpus vancampoi Zone. The transition between the zones is marked by a colour change in the sediments, the appearance of coals and organic-rich sediments above the transition, a decrease in numbers of vertebrate fossils, and the appearance of new pollen species. It is inferred that the observed changes reflect a climatic switch from dry to wetter conditions. At a large scale, palynomorph assemblages across this transition do not clearly indicate marked differences that can be attributed to the climatic change, with the exception of the appearance of Scollardia trapaformis and Mancicorpus gibbus in the assemblages. On a microscale of a few metres, there are changes in relative abundance of plant groups.

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